Graphis aphanes Mont. & Bosch

Kalb, Jutarat, Lücking, Robert & Kalb, Klaus, 2018, The lichen genera Allographa and Graphis (Ascomycota: Ostropales, Graphidaceae) in Thailand-eleven new species, forty-seven new records and a key to all one hundred and fifteen species so far recorded for the country, Phytotaxa 377 (1), pp. 448-450 : 448-450

Graphis aphanes Mont. & Bosch , in: Montagne (1856: 347).

= Graphis glaucocinerea Vain. (see also under excluded names).

Type:— JAVA. Ad corticem arborum, Teysman s.n. (L, not seen) .

Thallus corticolous, off-white, lead-gray to pale olive-green, smooth, dull. Lirellae prominent, with a complete thalline margin, short to elongate, straight, curved or sinuose, usually unbranched or very rarely branched, up to 10 mm long, labia convergent or seldom slightly divergent, entire, not or only indistinctly striate (illinata -morph), disc rarely visible from above, whitish pruinose. Exciple complete, brown to orange-brown, without carbonization; hymenium clear; ascospores (4–) 8/ascus, hyaline, transversely septate, 14–24-septate, 70–75 × 9–10 μm, 14–23-septate, 80–95 × 10–11 μm (Archer 2009), up to 100 μm long in the holotype (Makhija et al. 2005).

Chemistry: stictic acid (major), constictic acid (major to submajor), cryptostictic acid (minor), ± hypostictic acid (trace) (anal. K. Kalb).

Distribution and habitat:—SE-Asia, including Java (type locality), India, growing in a semi-evergreen forest in open places on roadside trees ( Awasthi 1991, Makhija et al. 2005), Japan ( Nakanishi et al. 2001), Australia (Archer 2009) and Thailand ( Nakanishi et al. 2001), Phitsanulok province: Nakhon Thai district, where it is growing on bark of an unidentified tree in a montane rainforest between 1250 and 1350 m.

Remarks:— Graphis aphanes was already reported for Thailand in Buaruang et al. (2017) as Hemithecium aphanes (Mont. & Bosch) M. Nakan. & Kashiw. , in: Nakanishi et al. (2003: 88). The species is very rare in Thailand (only one collection is known). It is characterized in having conspicuous off-white lirellae, long, multilocular ascospores and stictic acid with satellite substances. Makhija et al. (2005) reported also the occurrence of norstictic acid, but this acid was not found in five specimens from other regions in hb. Kalb tested by TLC.