Omphale sulciscuta (Thomson) Hansson, Christer & Shevtsova, Ekaterina, 2012

Hansson, Christer & Shevtsova, Ekaterina, 2012, Revision of the European species of Omphale Haliday (Hymenoptera, Chalcidoidea, Eulophidae), ZooKeys 232, pp. 1-157 : 103-107

Omphale sulciscuta (Thomson) comb. n. Figures 7, 9402-416497532

Derostenus (Holcopelte) sulciscuta Thomson, 1878:272. Holotype female in LUZM, examined.

Horismenus sulciscutus (Thomson), Schmiedeknecht (1909).

Holcopelte sulciscuta (Thomson), Graham (1959).

Holcopelte sulciscuta (Thomson), Bouček (1971).

Material.

Type material. Holotype female in LUZM (no type number). Additional material. 285♀ 6♂: Czech Republic 1♀ (BMNH), Denmark 1♀ 1♂ (LUZM), France 3♀ (BMNH), Hungary 43♀ 1♂ (BMNH, CH), Russia 12♀ (BMNH, CH, LUZM), Sweden 112♀ 1♂ (BMNH, CH, LUZM, NHRS), United Kingdom 113♀ 3♂ (BMNH).

Diagnosis.

Body strongly sclerotized and not collapsing after death (Figs 409, 413); head smooth and shiny (Figs 411, 412, 415, 416), frons black metallic with antennal scrobes joining below frontal suture in female (Figs 405, 411), in male bright metallic bluish green with antennal scrobes joining frontal suture separately (Figs 405, 415); occipital margin as a sharp carina (Fig. 412); notauli in female with posterior ½ as distinct and narrow grooves (Fig. 409); scutellum in anterior ⅔ with a distinct median groove (Figs 409, 413); propodeum with median and lateral longitudinal carinae (Figs 409, 413); petiole quadrangular with anterior part drawn out to a sharp edge that covers the propodeal nucha (Fig. 409); male gaster dark brown with a median white spot across tergites 1-3.

Description.

Female. Length of body 1.1-1.7 mm. Antenna with scape yellowish brown to pale brown with dorsal margin dark brown; pedicel and flagellum dark brown and shiny; pedicel + flagellum 1.3 × as long as distance between eyes; first flage llomere 1.0 × as long as second flagellomere, distinctly wider than remaining flagello-meres, 2.0 × as wide as second flagellomere (Fig. 410); flagellomeres 1-4 with setae confined to a basal whorl and with setae reaching beyond apex of flagellomere attached to; longitudinal sensilla on flagellomeres as long as flagellomere attached to; clava 1-segmented. Face black metallic with green tinges (Fig. 405), smooth (Fig. 411); cly-peus black metallic with green tinges, smooth, trapezoid to almost semicircular, 1.5 × as wide as high; gena black metallic; lower frons black metallic, smooth; interscrobal area smooth; antennal scrobes join below frontal suture; frontal suture V-shaped; upper frons and vertex black metallic, smooth (Fig. 412). Occipital margin with a sharp carina (Fig. 412).

Mesoscutum black metallic (Fig. 403) with engraved reticulation (Fig. 409), midlobe with two pairs of setae; notauli as distinct deep grooves in posterior ½. Scutellum black metallic (Fig. 403) with anteromedian ½ with engraved and weak reticulation, remainder with raised and strong reticulation (Fig. 409), with a distinct median groove in anterior ⅔; 1.2 × as long as wide, with anterior margin straight. Axillae black metallic (Fig. 403). Dorsellum black metallic (Fig. 403), tongue like (Fig. 409), smooth and with anterior ½ concave and sharply margined, 0.4 × as long as wide, and 0.3 × as long as length of median propodeum. Entire lateral mesosoma black metallic (Fig. 402); transepimeral sulcus weakly curved forwards. Propodeum black metallic (Fig. 403), smooth with a wide and shallow groove along anterior margin (Fig. 409), with a narrow median carina, laterally with a longitudinal carina half way between median carina and spiracular sulcus, posteromedian part slightly drawn out to form a short nucha that is delimited anteriorly by a transverse carina; propodeal callus with two setae. Foreleg with coxa dark brown (Fig. 402), femur pale brown, tibia yellowish brown to pale brown, tarsus dark brown; midleg with coxa dark brown, femur pale brown to dark brown, tibia and tarsus yellowish brown to pale brown, first tarsomere 0.2 × as long as length of tarsus; hind leg with very base of coxa dark brown to black, remaining coxa pale brown to dark brown, in some specimens the entire coxa is black, femur pale brown, tibia yellowish brown to pale brown, tarsus dark brown. Forewing transparent, veins and setae dark brown (Fig. 407); speculum closed; admarginal setae 8-12, arising mainly from wing membrane; radial cell setose; postmarginal vein 0.7 × as long as stigmal vein. Hind wing transparent, apex pointed (Fig. 407). Forewing WIP (Fig. 408) with apical ½ magenta with apical margin yellow, basal ½ with wide bands in blue, yellow and magenta.

Petiole black, quadratic and about as long as wide, to transverse, with anterior part drawn out to a sharp margin that covers propodeal nucha. Gaster dark brown to black and metallic, smooth, elongate and 1.3 × as long as length of mesosoma; 7th tergite 0.04 × as long as length of gaster.

Male. Length of body 1.2-1.7 mm. Features as in female except as follows. Antenna with scape dark brown with basal part yellowish white, pedicel and flagellum pale brown; pedicel + flagellum 1.9 × as long as distance between eyes; flagellomeres 1-4 with verticillate setae and with setae reaching beyond apex of flagellomere attached to (Fig. 414); clava 1-segmented. Face bright metallic bluish green (Fig. 406), smooth (Fig. 415); clypeus bright metallic bluish green; lower frons bright metallic bluish green; antennal scrobes join frontal suture separately; upper frons bright metallic bluish green; vertex black metallic with blue and green tinges.

Mesoscutum black metallic with purple tinges (Fig. 404), with engraved reticulation (Fig. 413). Scutellum black metallic with purple tinges (Fig. 404), predominantly with engraved reticulation (Fig. 413), posterior and lateral margins with raised and strong reticulation, with a distinct median groove in anterior ⅓; with median part of anterior margin protruding forwards. Axillae black metallic with purple tinges (Fig. 404). Dorsellum black metallic with purple tinges (Fig. 404), with longitudinal carinae (Fig. 413). Propodeum dark brown metallic (Fig. 404). Foreleg with coxa and femur dark brown, tibia and tarsus yellowish brown; midleg with coxa and femur dark brown, tibia and tarsus yellowish brown, first tarsomere 0.3 × as long as length of tarsus; hind leg with coxa and femur dark brown, femur tibia pale brown, tarsus yellowish white with 4th tarsomere dark brown, to entirely dark brown. Forewing admarginal setae 10, arising mainly from wing membrane.

Gaster dark brown with a median white spot across tergites 1-3, smooth, 0.9 –1.0× as long as length of mesosoma. Phallobase and aedeagus as in Fig. 497.

Host.

Unknown.

Distribution.

Armenian SSR ( Bouček and Askew 1968), Czech Republic ( Bouček 1957), Denmark (new record), France (new record), Germany ( Bouček and Askew 1968), Hungary ( Erdös 1956), Moldova ( Bouček 1965), Russia (new record), Sweden ( Thomson 1878); United Kingdom ( Graham 1959) (Fig. 532).

Remarks.

The male is described here for the first time. Similar to several other Omphale species the sex ratio in the material at hand is skewed with considerably more females than males.

Unplaced species