Ptecticus

Rozkošný, Rudolf & Hauser, Martin, 2009, Species groups of Oriental Ptecticus Loew including descriptions of ten new species with a revised identification key to the Oriental species (Diptera: Stratiomyidae), Zootaxa 2034, pp. 1-30: 25-28

publication ID

http://doi.org/ 10.5281/zenodo.186331

persistent identifier

http://treatment.plazi.org/id/644E87FB-2571-FFE9-FF5B-6F2E2BDCF926

treatment provided by

Plazi

scientific name

Ptecticus
status

 

Key to the Oriental species of Ptecticus  

Compared with the last version of the identification key to the Oriental Ptecticus   species ( Rozkošný & Kovac 2003) the following changes are made: (1) ten new species described above, and two additional species, viz., P. thailandicus Rozkošný & Courtney, 2005   and P. kovaci Hauser & Rozkošný, 2008   are included; (2) some recently confirmed synonyms are incorporated – P. brunettii Rozkošný & Kovac, 1996   is a synonym of P. brevipennis ( Rondani, 1875)   (see Mason & Rozkošný 2005 a), P. repensans ( Walker, 1859)   is a synonym of the predominantly Australasian P. complens ( Walker, 1858)   (see Mason & Rozkošný 2005 b), P. minimus Rozkošný & Kovac, 1997   is a synononym of P. shirakii Nagatomi, 1975   , P. tenebrifer ( Walker, 1849)   is a synonym of P. japonicus ( Thunberg, 1789)   and P. wulpii Brunetti, 1907   is a synonym of P. melanurus ( Walker, 1848)   (comments on the two latter synonyms see above); (3) P. mirabilis Rozkošný & Kovac, 2003   was ommited in the last key but it is actually a synonym of P. anneliesae Lindner, 1935   (see Hauser & Rozkošný 2008); (4) P. sinchangensis Ôuchi, 1938   was not included in the new key owing to its Palaearctic distribution and the occurrence of the Australasian P. rogans ( Walker, 1859)   needs revision because records of this Australasian species from India and the Philippines ( Brunetti 1907) seem to be problematic (cf. Brunetti 1923); on the contrary, the Australasian P. ferrugineus ( Doleschall, 1859)   probably also occurs in Sulawesi and is thus included; (5) references to the published illustrations of distinguishing characters and basic distributional data are added.

1 Median occipital (postvertical) sclerite yellow, contrasting with rest of occiput ......................................................... 2

- Median occipital sclerite black or rarely brown and thus not or barely distinct from black postocular part of occiput .................................................................................................................................................................................... 22

2 Scutum with three well defined, broad, dark longitudinal vittae, rarely blackish only on sides along notopleuron (his- trio -group, part) ............................................................................................................................................................ 3

- Scutum yellowish brown to brown, without definite pattern or predominantly black with metallic reflections ........ 8

3 Upper frons shining black (male terminalia and abdomen: figs 16–20 in original description) ( Thailand) .................. .................................................................................................................. P   . pangmapensis Rozkošný & Kovac, 2003

- Upper frons yellow to yellowish brown ...................................................................................................................... 4

4 Scutum darkened only laterally along notopleuron (body including male terminalia: figs 10–17 in Mason & Roz- košný 2005 a) ( Thailand and Malaysia) ....................................................................... P. brevipennis ( Rondani, 1875)  

- Scutum with three broad dark longitudinal vittae, only narrow yellow lines among them yellow ............................. 5

5 Katatergite and upper part of mediotergite yellow (body parts including male terminalia figs 11–19 in Rozkošný & Kovac, 1994) ( Laos, Malaysia)...................................................................... P. malayensis Rozkošný & Kovac, 1994  

- Katatergite and mediotergite black ............................................................................................................................... 6

6 Hind basitarsus bicoloured in female, black and yellow (abdomen: fig. 1 in original description; male unknown ( Sri Lanka) ....................................................................................................... P. pseudohistrio Rozkošný & Hauser, 2001  

- Hind basitarsus uniformly black in female, black or bicoloured in male ................................................................... 7

7. Abdominal tergites 4 and 5 extensively black, usually without posterior yellow band (abdomen and male terminalia:

figs 6–9 in Rozkošný & De Jong 2001) ( Indonesia: Java) ................................................ P. histrio   De Meijere, 1933 - Abdominal tergites 4 and 5 with posterior yellow band (male terminalia figs 20–23 in original description) ( Laos, Malaysia, Singapore, Thailand) ........................................................................ P. proximus Rozkošný & Kovac, 1996  

8 Upper frons shining black ........................................................................................................................................... 9

- Upper frons yellow or yellowish brown .................................................................................................................... 12

9. Wing intensively brownish infumated or with darkened apex ................................................................................ 10

- Wing without distinct infumation .............................................................................................................................. 11

10 Scutum black with metallic green reflections; wing hyaline, only apex blackish; hind leg dark, only trochanter, knee and tarsomeres 2–3 pale yellow to white; male unknown ( Philippines)........ P. fumipennis Rozkošný & Kovac, 2003  

- Scutum yellowish brown; whole wing intensively brownish infumated; hind leg black including trochanter, basal half of tibia pale yellow (female terminalia: Figs 28–30 View FIGURES 28 – 30 ), male unknown (East Malaysia) ....... P. infuscatus   sp. nov.

11 Apical three tarsomeres of hind leg snow-white (male terminalia: figs 1–4 in original description) ( Indonesia: Sulawesi, East Malaysia, Singapore) .................................................................... P   . bifidus Rozkošný & Kovac, 1998

- Apical three tarsomeres of hind leg black (male terminalia: figs 5–7 in original description) (East Malaysia) ............ ......................................................................................................................... P. bilobatus Rozkošný & Kovac, 1998  

12. Hind femur blackish in apical half; male cerci unusually elongated ( tenebrifer   group, part) .................................... 13

- Hind femur uniformly ochre yellow or darkened in basal half; cerci not unusually elongated ............................... 14

13 Area above fore coxa black and with some black hairs, anepimeron blackish, male unknown ( Laos) ......................... ................................................................................................................................................ P. vitalisi Brunetti, 1924  

- Area above fore coxa at most brown, without black hairs, anepimeron yellow (male terminalia and abdomen: figs 1 –2, 9 in original description) ( Laos, East Malaysia, Thailand) ......................... P. erectus Rozkošný & Kovac, 2000  

14 Wing at least partly infuscated; male cerci mostly short and broad; proctiger characteristically concave ventrally ( aurifer   group) ........................................................................................................................................................... 15

- Wing almost hyaline, at most very finely and uniformly yellowish brown tinged; male cerci usually slender and pointed; proctiger not concave ventrally..................................................................................................................... 20

15 Greater part of pleura, abdominal venter and basal half of hind femur black .......................................................... 16

- Pleura, at least basal half of abdominal venter and entire hind femur ochre yellow ................................................. 17

16 Wing uniformly brown infuscated; male gonostylus simply pointed apically (male terminalia as in P. aurifer   ) ( India, Myanmar) ...................................................................................................................... P   . aurobrunneus Brunetti, 1920

- Wing orange yellow, darkened on apex and along posterior margin; male gonostylus bicuspidate apically (male terminalia: Figs 22–24) ( Laos) ........................................................................................................ P. subaurifer   sp. nov.

17 Male cerci slender, gonostyli shifted to middle, also slender and narrow (abdomen and male terminalia: figs 18–21 in Rozkošný & De Jong 2001); tip of female abdomen beginning at segment 6 reddish yellow ( China, India, Indonesia: Sumatra, Laos, Nepal, Thailand) ................................................................................. P   . kerteszi De Meijere, 1924

- Male cerci broad, nearly triangular, gonostyli broader and in normal position ( Figs 19–21 View FIGURES 19 – 21 ) .................................. 18

18 Wing uniformly infuscated, not bicoloured, tip of female abdomen beginning at segment 5 black (East Malaysia) ... ................................................................................................................................................ P. sarawakensis   sp. nov.

- Wing bicoloured orange at the base and blackish at the tip ....................................................................................... 19

19 Darkening of wing apex reaching discal cell or nearly so; basal 3–4 abdominal segments usually ochre yellow (body parts including. male terminalia: figs 18–25 in Mason & Rozkošný 2005 a) (from India and China to Singapore and Laos) ................................................................................................................................ P. melanurus ( Walker, 1848)  

- Darkening of wing apex reaching usually only middle or slightly more of distance between discal cell and wing margin; basal abdominal segments usually with extensive dark pattern or predominantly brown (male terminalia: figs. 8–9 in Nagatomi 1975) (East Palaearctic and Oriental) ...................................................... P. aurifer ( Walker, 1854)  

20 Hind basitarsus uniformly yellowish brown; male terminalia asymmetrical (especially aedeagal complex, Figs 4–7 View FIGURES 4 – 7 ) ( histrio   group, part) ( Laos)....................................................................................................... P. bannapensis   sp. nov.

- Hind basitarsus contrastingly brown or black at least in basal half; male terminalia symmetrical ( cingulatus   group, part) ........................................................................................................................................................................... 21

21 Abdominal transverse bands reduced, barely occupying middle third of tergites 2–4; male gonostylus trilobate (abdomen and male terminalia: figs 22–25 in original description) ( Philippines) ......................................................... .................................................................................................................. P   . philippinensis Rozkošný & Kovac, 2003

- Abdominal transverse bands not reduced, much broader than half length of tergites 2–4; male gonostylus simply pointed apically (abdomen and male terminalia: figs 26–30 in original description) ( Singapore) ................................ ................................................................................................................... P. temasekianus Rozkošný & Kovac, 2003  

22 Hind legs completely black; male cerci unusually long ( tenebrifer   group, part) ..................................................... 23

- Hind legs at least partly yellow; male cerci not unusually long ................................................................................ 24

23 Abdominal tergite 2 with at least 3 black spots; abdomen spindle-shaped (body including male terminalia: figs 12–19 in Mason & Rozkošný 2005 b) ( Indonesia: Sulawesi and Philippines) ..................... P. remeans   ( Walker, 1860)

- Abdominal tergite 2 translucent whitish, with a small medial spot; abdomen more spatulate (male terminalia figs

14–15 in Nagatomi 1975 under P. tenebrifer   ) (East Palaearctic, China, Taiwan......... P. japonicus ( Thunberg, 1789)   24 Postcubitus (CuP) distinct, pigmented; M 3 distinctly sinuate ( Figs 38 View FIGURES 34 – 41. 34 – 35 and 48 View FIGURES 42 – 49. 42 – 43 ).......................................................... 25

- Postcubitus (CuP) developed at most as hyaline wing fold; M 3 straight, parallel to M 2 ( Figs 36 View FIGURES 34 – 41. 34 – 35 , 42, 44 View FIGURES 42 – 49. 42 – 43 ).................. 37

25 Scutum with three broad dark longitudinal vittae ( tricolor   group) ........................................................................... 26

- Scutum uniformly yellow or with another pattern ( cingulatus   group, part)............................................................... 28

26 Hind femur yellow, at most its basal third brownish; abdominal tergites 1 and 2 without dark pattern ( Singapore) .... ........................................................................................................................................... P. intensivus Brunetti, 1927  

- Apical half (or somewhat less) of hind femur black; abdominal tergites 1 and 2 with black pattern ........................ 27

27 Abdomen black, tergites 2–4 usually red; hind tarsus completely black (body including male terminalia: figs 24–30 in Rozkošný & Kovac, 1996) ( India, Indonesia: Java, Sumatra; Thailand) ........ P. tricolor   Wulp in de Meijere, 1904

- Abdomen yellow, with extensive black pattern; hind tarsus broadly white in middle (male terminalia: figs 8–12 in original description) ( Laos, Thailand) ............................................................. P. siamensis Rozkošný & Kovac, 1998  

28 Scutum predominantly metallic blue (female terminalia: Figs 25–27 View FIGURES 25 – 27 ); male unknown .................. P   . elegans sp. nov.

- Scutum not metallic blue .......................................................................................................................................... 29

29 Hind femur uniformly yellow or yellowish brown.................................................................................................... 30

- Hind femur with black pattern .................................................................................................................................... 35

30 Epandrium longer than broad ................................................................................................................................... 31

- Epandrium barely longer than broad ......................................................................................................................... 34

31 Cerci not extending beyond apex of proctiger, gonostylus with hook-like apical projection (figs 5–9 in Hauser & Rozkošný 2008) ( Indonesia: Sulawesi) ............................................................................. P   . anneliesae Lindner, 1935

- Cerci distinctly extending beyond apex of proctiger, gonostylus without apical projection ( Figs 7–10 View FIGURES 4 – 7 View FIGURES 8 – 10 ) ................. 32

32 Posterior margin of genital capsule with a high medial process, aedeagal complex pointed apically ( Figs 11–14 View FIGURES 11 – 14 ) ( India) ............................................................................................................................................... P. indicus   sp. nov.

- Posterior margin of genital capsule without or with only very low medial process, aedeagal complex tube-like, rounded apically in dorsal view ................................................................................................................................. 33

33 Gonostylus longer than half length of genital capsule, dilated and rounded distally; apical segment of female cerci much longer (figs 15–18 in Rozkošný & De Jong 2003) ( Indonesia: Sulawesi and Australasian)................................ ................................................................................................................................... P   . ferrugineus ( Doleschall, 1859)

- Gonostylus barely as long as a quarter of genital capsule length, with a short inner subapical tip; apical segment of female cerci very short, barely longer than broad (male and female terminalia in figs 1–8 in original description) ( Thailand) ................................................................................................. P   . thailandicus Rozkošný & Courtney, 2005

34 Distal 4 tarsomeres on fore and mid legs dark and densely black haired; male epandrium almost round and cerci small (body including male terminalia: figs 15–19 in original description) ( Malaysia, Thailand) ............................... .................................................................................................................. P. flavifemoratus   Rozkošný & Kovac, 1996

- Distal 4 tarsomeres on fore and mid legs yellow, at most 4 th and and 5 th tarsomeres slightly darkened and with black hairs; epandrium subquadrate, cerci larger, extending beyond proctiger (figs 2–4 in original description) ( Philippines) .................................................................................................................... P. kovaci Hauser & Rozkošný 2008  

35 Scutum with diffuse black pattern: broad notopleural vittae and a linear median vitta that is triangularly expanded on both ends (body parts including male terminalia: figs 12–17 in Rozkošný & De Jong 2001) ...................................... ............................................................................................................................... P. kambangensis De Meijere, 1914  

- Scutum uniformly yellowish brown .......................................................................................................................... 36

36 Abdomen with well defined black transverse bands (head, abdomen and male terminalia in Rozkošný & Kovac 1996) (from India to Taiwan and East Malaysia) .................................................................. P. cingulatus Loew, 1855  

- Abdominal transverse bands often reduced, sometimes barely visible below dense yellow hairs or replaced by darkened hair patches (body including male terminalia: in Mason & Rozkošný 2005 b) ( Indonesia: Sulawesi; Philippines and Australasian)................................................................................................................ P. complens ( Walker, 1858)  

37 Vein R 2 + 3 arising far before anterior crossvein, very short or indistinct ( Figs 44–45 View FIGURES 42 – 49. 42 – 43 ); relatively small species (6.3–7.2 mm) (body including male terminalia: figs 1–9 in Rozkošný & Kovac 1997 under P. minimus   ) ( Laos, Malaysia, Thailand, Ryukyu Is.) ( shirakii   group) ............................................................................... P. shirakii Nagatomi, 1975  

- R 2 + 3 arising close to anterior crossvein, not confluent with R 1 ( Figs 36 –37 View FIGURES 34 – 41. 34 – 35 , 42– 43 View FIGURES 42 – 49. 42 – 43 ); usually larger species ............. 38

38 Scutum at least partly black with metallic blue reflections; dark area never confined only to median prescutellar area and scutellum ( longipennis   group, part) .................................................................................................................... 39

- Scutum uniformly yellow to yellowish brown; if partly metallic green, then dark area confined to median prescutellar area and scutellum ................................................................................................................................................ 42

39 Wing apex distinctly infumated; wings conspicously elongate ................................................................................ 40

- Wing apex not infumated; wings not conspicuously elongate................................................................................... 41

40 Medial process of male genital capsule very slender, formed of two rodlike and pointed projections (visible even in situ; heads and male terminalia: figs 1–6 in Rozkošný & Kovac, 1994); abdominal pattern very variable though usu-

ally consisting of transverse bands (from India to the Philippines, Indonesia and East Malaysia) ................................ ............................................................................................................................... P. longipennis ( Wiedemann, 1824)   - Medial process of synsternite of another shape; yellow abdominal pattern confined to anterior and posterior corners of tergites (abdomen and male terminalia: figs 9–12 in Rozkošný 2002) ( Indonesia: Sumatra).................................... ........................................................................................................................................ P. violaceus   ( Enderlein, 1914)

41 Hind femur completely yellow; male unknown ( India, Laos, Malaysia) ........................ P. assamensis Brunetti, 1923  

- Hind femur with a dark medial ring and a broad ventral streak in basal half (abdomen and male terminalia figs 1–5 in Rozkošný & De Jong 2001) ( Indonesia: Sumatra, Malaysia, Thailand) ............... P. aeneithorax De Meijere, 1919  

42 Wing apex infumated ( Fig. 42 View FIGURES 42 – 49. 42 – 43 ), fore and mid tarsomeres not annulated ( longipennis   group, part)........................... 43

- Wing apex not infumated ( Fig. 36 View FIGURES 34 – 41. 34 – 35 ); if rarely infumated than fore and mid tarsomeres markedly annulated ( australis   group)......................................................................................................................................................................... 44

43 Scutum and scutellum yellowish brown, with bronze to violet reflections, distinctly darker than yellow pleura; large species (14.2 mm) (male terminalia: figs. 6–11 in Rozkošný & Kovac 2003) ( Nepal and Laos) ................................. ....................................................................................................................... P. longispinus   Rozkošný & Kovac, 2003

- Entire thorax uniformly ochre yellow; small species (7.5 mm) (male terminalia: figs 5–8 in Rozkošný 2002) ( Indonesia: Sumatra, Java; East Malaysia) ............................................................................ P. sumatranus Enderlein, 1914  

44 Fore and mid tarsomeres markedly annulated with black and yellow (fore tarsus, abdomen and male terminalia: fig. 1–5 in original decription) ( Malaysia) ............................................................. P. annulipes Rozkošný & Kovac, 2003  

- Fore and mid tarsomeres not annulated .................................................................................................................... 45

45 Lower half of frontal callus markedly darkened (body including male terminalia: figs. 28–32 in Rozkošný & De Jong 2001) (from India to Japan: Ryukyu Is., Taiwan and Indonesia: Java) ................. P. vulpianus   ( Enderlein, 1914)

- Frontal callus completely pale ................................................................................................................................... 46

46 Males .......................................................................................................................................................................... 47

- Females ...................................................................................................................................................................... 52

47 Surstyli not developed ( Fig. 11 View FIGURES 11 – 14 ) ................................................................................................................................ 48

- Surstyli well developed (visible even in situ) ( Figs 1 View FIGURES 1 – 3 , 4 View FIGURES 4 – 7 ) ........................................................................................... 49

48 Gonocoxites (posterolateral parts of genital capsule) without conspicuous inner dorsal prominences but with slender inner ventral processes (see figs 13–15 in original description) ( Thailand, Malaysia, Indonesia: Sumatra) ................ ...................................................................................................................... P. subaustralis   Rozkošný & Kovac, 1998

- Gonocoxites with bilobed inner dorsal prominence; inner ventral processes replaced by a small bilobed medial projection (male terminalia: Figs 15–18 View FIGURES 15 – 18 ) ( Laos).................................................................................... P. kubani   sp. nov.

49 Medial process at posterior margin of genital capsule large....................................................................................... 50

- Medial process at posterior margin of genital capsule indistinct or small ( Fig. 3 View FIGURES 1 – 3 )..................................................... 51

50 Medial process of genital capsule subquadrate, with a deep medial incision, aedeagal complex massive, bipartite (figs 6 and 7 in Rozkošný & Hauser 1998 under P. ceylonicus   ) (from India and Sri Lanka to Vietnam and Cambodia) .................................................................................................................... P. srilankai Rozkošný & Hauser, 2001  

- Medial process of genital capsule consisting of two slender, slightly convergent processes (male terminalia: Figs 8–10 View FIGURES 8 – 10 ) (South China) .................................................................................................................. P. fukienensis   sp. nov.

51 Surstylus markedly dilated in apical half (abdomen and male terminalia: figs 8–12 in Rozkošný & Hauser 1998) .... .............................................................................................................................................. P. australis Schiner, 1868  

- Surstylus rod-like, pointed apically (male terminalia: Fig. 1–3 View FIGURES 1 – 3 ) ( Philippines) ................ P. artocarpophilus   sp. nov.

52 Basal half of hind basitarsus black .......................................................................................................................... 53

- Only extreme base of hind basitarsus black .............................................................................................................. 54

53 Median prescutellar area and greater part of scutellum dark and metallic green (female terminalia: Figs 31–33 View FIGURES 31 – 33 ), male unknown ( Nepal).................................................................................................................. P. semimetallicus   sp. nov.

- Scutum and scutellum uniformly yellow or yellowish brown (from India and Sri Lanka to Japan: Ryukyu Is; Taiwan and Indonesia: Krakatau Is.) ................................................................................................ P. australis Schiner, 1868  

54 Visible parts of tergites 6 and 7 wholly yellow (from India and Sri Lanka to Vietnam and Cambodia)........................ .......................................................................................................................... P. srilankai Rozkošný & Hauser, 2001  

- Visible parts of tergites 6 and 7 with black pattern ( Thailand, Malaysia, Indonesia: Sumatra) ..................................... ...................................................................................................................... P. subaustralis   Rozkošný & Kovac, 1998