Tarocystis responsivus ( Hedley, 1912 )

Hyman, Isabel T. & Ponder, Winston F., 2010, A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462, Zootaxa 2462 (1), pp. 1-148 : 78-79

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https://doi.org/ 10.11646/zootaxa.2462.1.1

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Tarocystis responsivus ( Hedley, 1912 )


Tarocystis responsivus ( Hedley, 1912)

Figures 10E View FIGURE 10 , 11E View FIGURE 11 , 16C, G View FIGURE 16 , 24F View FIGURE 24 , 25J–L View FIGURE 25 , 26F View FIGURE 26 , 27F View FIGURE 27 , 28F View FIGURE 28

Microcystis responsivus Hedley, 1912: 262 .

Tarocystis responsivus: Iredale, 1937c: 4 ; Smith, 1992: 243.

Material examined. Queensland: Syntypes: AM C104308 (one shell measured, photographed), W of Gladstone, Port Curtis (23º51' S, 151º15' E), on ground under leaves in bottle tree scrub, Aug. 1908, S.W. Jackson GoogleMaps .

Other material: AM C425327 (two specimens dissected, radulae examined), Eurimbula State Forest , S of Gladstone (24º11' S, 151º50' E), on ground, 7 May 1975, J.B. Burch, W.F. Ponder, P.H. Colman GoogleMaps .

Description. External morphology: Shell ( Figures 16C, G View FIGURE 16 , 26F View FIGURE 26 , 27F View FIGURE 27 , 28F View FIGURE 28 ) 4 whorls, pale orange-brown; shape and sculpture as for genus. Animal white. Mantle laps and lobes as for genus. Caudal horn large; caudal foss vertical slit in tail.

Mantle cavity ( Figure 10E View FIGURE 10 ) and digestive system: As for genus.

Genital system ( Figure 24F View FIGURE 24 ): As for genus. Penis moderately long; epiphallus enters penis through simple pore; penis internally flat, at least one longitudinal penis pilaster present. Epiphallus shorter than penis, internally with longitudinal pilasters. Epiphallic retractor caecum absent. Flagellum and lime-sac absent.

Radula ( Figure 25J–L View FIGURE 25 ): As for genus. Radular formula ( × 79 rows.

Range and habitat. Tarocystis responsivus is found from Mt. Nebo, W of Brisbane, to Crocodile Creek, SW of Rockhampton, SE Queensland, in scrub, open sclerophyll forest and microphyll and notophyll vine thickets. Specimens are usually found on the ground in leaf litter and under rocks and logs.

Remarks. The genitalia of Tarocystis fulva figured in Odhner (1917) shows a long, slender flagellum, which is absent in T. responsivus . However, the long slender vagina and bursa copulatrix (mistakenly labelled as a dart sac) are both present in T. responsivus . The bursa copulatrix of T. fulva was described as containing a ‘chitinous chord’, probably a spermatophore being broken down. No spermatophores were observed in T. responsivus ; however, the lack of a flagellum in this species may indicate that a hard-walled spermatophore is not formed.

The other two species in this genus, Tarocystis antiqua and T. fulva , are both found further north (their type localities are Chillagoe Caves and Atherton Tableland respectively).


Australian Museum














Tarocystis responsivus ( Hedley, 1912 )

Hyman, Isabel T. & Ponder, Winston F. 2010

Tarocystis responsivus: Iredale, 1937c: 4

Smith, B. J. 1992: 243
Iredale, T. 1937: 4

Microcystis responsivus

Hedley, C. 1912: 262