Euconnus (Glabriconnus) andranomenaus, Jałoszyński, 2016

Jałoszyński, Paweł, 2016, Taxonomy of ' Euconnus complex'. Part VII. Glabriconnus, a new subgenus of Euconnus from Madagascar (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4084 (4) : -

Euconnus (Glabriconnus) andranomenaus sp. n.

( Figs 16–19 , 23–25 , 30–31 , 38 )

Type material. Holotype (MADAGASCAR): ♂, two labels: "ADR/ Feb.2013/07 MADAGASCAR / ANDRANOMENA Sp. Res. 16 m, / „Circuit Andranohazo”, 2.ii. 2013 / S20º0.9'13.1''E44º30'29.1'' / sifting litter; Winkler app. extr.; L.S. / Rahanitriniaina & E.M. Rabotoson lgt." [white, printed], " EUCONNUS / (GLABRICONNUS) / andranomenaus m. /det. P. Jałoszyński, 2015 / HOLOTYPUS" [red, printed] (MMBC). Paratypes (4 exx.): 2 ♂♂, same data as holotype; 1 ♂, "ADR/ Feb.2013/06 MADAGASCAR / ANDRANOMENA Sp. Res. 29 m, / „Circuit Andranohazo”, 2.ii. 2013 / S20º0.9'18.6''E44º30'20.9'' / sifting litter; Winkler app. extr.; L.S. / Rahanitriniaina & E.M. Rabotoson lgt."; 1 ♂, "ADR/ Feb.2013/15 MADAGASCAR / ANDRANOMENA Sp. Res. 30 m, / „Circuit Trois Lacs”, 3.ii. 2013 / S20º0.8'58.6''E44º31'09.6'' / sifting litter; Winkler app. extr.; L.S. / Rahanitriniaina & E.M. Rabotoson lgt." (MMBC, cPJ).

Diagnosis. Small species, BL <2 mm; frons with two rounded longitudinal carinae converging posteriorly; pronotum with two parallel lateral hypomeral grooves at each side; elytra oval, similarly narrowing posteriorly and anteriorly; aedeagus only slightly more than twice as long as broad, broadest in proximal half.

Description. Body of male ( Figs 16–17 ) strongly convex, elongate with deep constriction between head and pronotum and between pronotum and elytra, appendages long; BL 1.86–1.93 mm (mean 1.90 mm); cuticle glossy and slightly opalescent, reddish-brown, legs and antennae slightly lighter, palps and tarsi light brown; setae yellowish, bristles nearly as dark as cuticle.

Head ( Figs 16, 18 ) in dorsal view approximately subpentagonal, broadest at eyes, HL 0.34–0.35 mm (mean 0.35 mm), HW 0.38 mm; vertex and frons confluent, frons and anterior portion of vertex with pair of longitudinal median carinae, which are rounded and weakly elevated but distinct and distinctly converging but not connecting posteriorly, area between carinae distinctly impressed; supraantennal tubercles weakly raised but distinct; tempora about twice as long as eyes in dorsal view; eyes moderately large and moderately convex, coarsely faceted; head in lateral view ( Figs 17, 19 ) distinctly flattened and with vertex slightly bulging posterodorsally. Punctures on vertex and frons very fine and sparse, inconspicuous; setae on vertex and posterior portion of frons indiscernible, only frons between supraantennal tubercles with several moderately long and suberect setae; genae and postgeneae, but not tempora, with thick bristles. Antennae moderately long, AnL 0.70–0.75 mm (mean 0.73 mm); antennomeres I– VII distinctly elongate, VIII about as long as broad, IX–X each distinctly transverse, XI not broader than X, much shorter than IX–X together, only about 1.4× as long as broad, with subconical and indistinctly pointed apex.

Pronotum in dorsal view ( Fig. 16 ) bell-shaped, broadest distinctly in anterior third, slightly variable in proportions, in most specimens slightly broader than long, in one as broad as long, PL 0.50–0.53 mm (mean 0.51 mm), PW 0.51–0.53 mm (mean 0.52 mm); anterior margin weakly arcuate, anterior corners weakly marked, obtuse-angled and blunt; lateral margins strongly rounded in anterior half and distinctly, narrowly and shallowly constricted just in front of posterior corners, which are slightly obtuse-angled or nearly right-angled, blunt; posterior margin slightly bisinuate; base with one distinct pair of deep lateral pits; sublateral carinae very indistinct; each side of pronotum with two parallel hypomeral grooves extending along nearly entire length of pronotum, dorsal groove in posterior half partly visible in dorsal view. Punctures on pronotal disc not visible at magnifications up to 80×; vestiture composed of very sparse but distinct, moderately long and suberect setae distributed on pronotal disc and dense bristles along its anterior margin, also area between lateral hypomeral grooves covered with bristles ( Fig. 19 ).

Elytra ( Figs 16–17 ) oval, broadest near middle, EL 1.03–1.08 mm (mean 1.05 mm), EW 0.80–0.85 mm (mean 0.82 mm), EI 1.26–1.31; elytral base slightly broader than pronotal base, lacking impressions and humeral calli. Punctures on elytral disc as fine as those on pronotum; setae very sparse and short, nearly recumbent. Hind wings absent.

Legs ( Figs 16–18 , 23 ) moderately long and slender; protibiae modified, with distal third or fourth curved ventrally and with ventral apical cuticular spur-like projection forming obtuse angle with ventral tibial margin, which bears tuft of setae just behind projection ( Fig. 23 ).

Metaventrite with small distinct median tubercle near posterior margin.

Aedeagus ( Figs. 24–25 , 30–31 ) moderately elongate, AeL 0.55 mm, in ventral view broadest near middle of capsular portion of median lobe with abruptly and strongly narrowing, nearly rod-like apical part, its apex distinctly broadened; in lateral view median lobe with apical portion slightly bent dorsally and apex curved ventrally; endophallus strongly asymmetrical, with two subapical bunches of dense elongate needle-like sclerites pointing distally and laterally; parameres broad, each with several apical and subapical setae not reaching apex of median lobe.

Female. Unknown or indistinguishable from E. februarius (see Remarks).

Etymology. Locotypical, after Andranomena Reserve.

Distribution. Central-western Madagascar ( Fig. 38 ).

Remarks. This species cannot be distinguished from E. februarius sp. n. on the basis of external characters; only examination of aedeagus ensures correct determination. For this reason numerous females collected together with males of these two species cannot be unambiguously identified and the descriptions are based on males only.

Figures

FIGURE 38. Distribution of Euconnus (Glabriconnus) spp. in Madagascar.

FIGURES 16 – 19. Euconnus (Glabriconnus) andranomenaus sp. n., male. Dorsal (16) and lateral (17) habitus; head in dorsal view (18); head and prothorax in lateral view (arrows showing hypomeral grooves) (19).

FIGURES 20 – 27. Euconnus (Glabriconnus) banari sp. n. (20 – 22), Euconnus (Glabriconnus) andranomenaus sp. n. (23 – 25) and Euconnus (Glabriconnus) februarius sp. n. (26 – 27). Left protibia of male in mesal (inner) view (20, 23); aedeagus in ventral (21, 24, 26) and lateral (22, 25, 27) views.

FIGURES 28 – 33. Euconnus (Glabriconnus) banari sp. n. (28 – 29), Euconnus (Glabriconnus) andranomenaus sp. n. (30 – 31) and Euconnus (Glabriconnus) februarius sp. n. (32 – 33). Aedeagus in ventral (28, 30, 32) and lateral (29, 31, 33) views.