Kaliella punctata Vermeulen, Liew & Schilthuizen

Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno, 2015, Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species, ZooKeys 531, pp. 1-139 : 74-76

Taxon classification Animalia Stylommatophora Euconulidae

Kaliella punctata Vermeulen, Liew & Schilthuizen sp. n. Figure 64, 67

Kaliella punctata nomen nudum, Clements et al. 2008: 2762; Schilthuizen et al. 2013: online supplementary data.

Coneuplecta angulata auct., Schilthuizen et al. 2002: 256-257; Schilthuizen et al. 2002b: 37, 41-42; Schilthuizen and Vermeulen 2003: 96; Clements et al. 2008: 2762.

Holotype. Malaysia, Sabah, Interior Province , Sepulut River valley , Gua Pungiton ( RMNH.5003925 ).

Examined material from Sabah.

Interior Province. Crocker Range N.P., Gua Laing c. 12 km North of Keningau (leg. J.J. Vermeulen, V 1108; leg. M. Schilthuizen, BOR/MOL 821). Gunung Trusmadi slopes, Gua Loloposon (leg. J.J. Vermeulen, V 13235). Pinangah valley, Batu Urun (= Bukit Sinobang) (leg. J.J. Vermeulen, V 8005). Pun Batu c. 30 km West of Sepulut (leg. J.J. Vermeulen, V 1277). Sepulut valley, Batu Punggul (leg. J.J. Vermeulen, V 1987; leg. M. Schilthuizen, BOR/MOL 788; leg. T.S. Liew, M. Schilthuizen & S. Chiba, BOR/MOL 4325); Batu Temurung (leg. J.J. Vermeulen, V 8052; leg. M. Schilthuizen, BOR/MOL 819); Bukit Tinagas, East end of Batu Punggul limestone (leg. J.J. Vermeulen & M. Schilthuizen, V 7632); Gua Pungiton (leg. J.J. Vermeulen & M. Schilthuizen, V 7829, BOR/MOL 786); Gua Sanaron (leg. J.J. Vermeulen & M. Schilthuizen, V 7669; leg. M. Schilthuizen, BOR/MOL 794, BOR/MOL 806). Sandakan Province. Kinabatangan valley, Batu Keruak 2 near Sukau (leg. J.J. Vermeulen & M. Schilthuizen, V 9796, BOR/MOL 2364; leg. M. Salverda & H. van Oosten, BOR/MOL 2367); Batu Materis (leg. T.S. Liew & B. Elahan, BOR/MOL 2119; leg. M. Schilthuizen, BOR/MOL 2368); Batu Tai (not Bod Tai) near Gomantong (leg. J.J. Vermeulen & M. Schilthuizen, V 9605, BOR/MOL 2365; leg. T.S. Liew & B. Elahan, BOR/MOL 1918); Bod Tai (leg. M. Schilthuizen, BOR/MOL 2366); Batu Tomanggong Besar (leg. M. Schilthuizen, BOR/MOL 795); Batu Tomanggong Kecil (leg. J.J. Vermeulen & M. Schilthuizen, V 9694; leg. T.S. Liew & B. Elahan, BOR/MOL 2029, BOR/MOL 2059); Batu Tulug (Batu Putih) along road Lahad Datu-Sandakan, North of bridge over Kinabatangan River (leg. J.J. Vermeulen & H. Duistermaat, V 1485; leg. M. Schilthuizen, BOR/MOL 796); Gomantong Hill 30 km South of Sandakan (leg. J.J. Vermeulen & H. Duistermaat, V 1616; leg. A. van Til, BOR/MOL 3314; leg. T.S. Liew & J.P. King, BOR/MOL 3666); Tandu Batu (leg. J.J. Vermeulen & M. Schilthuizen, V 9629). Segama Valley, North end of limestone ridge on East bank Tabin River (leg. J.J. Vermeulen & M. Schilthuizen, V 7783, BOR/MOL 791), Tabin Wildlife Reserve (leg. T. Kimsin & H.N. Chai, BOR/MOL 800; leg. M. Schilthuizen, BOR/MOL 822). Tawau Province. Batu Baturong c. 50 km W.S.W. of Lahad Datu (leg. J.J. Vermeulen & H. Duistermaat, V 1859). Segama valley, hill N.W. of crossing road Sandakan-Lahad Datu with the Segama River (leg. J.J. Vermeulen & H. Duistermaat, V 1682); Danum Valley (leg. M. Schilthuizen, BOR/MOL 797, BOR/MOL 798, BOR/MOL 811; leg. H.A. Rutjes, BOR/MOL 3618, BOR/MOL 792; leg. UMS students, BOR/MOL 787; leg. T. Kimsin & H.N. Chai, BOR/MOL 799); ‘Kirk’s Cave’ 8 km North of Lahad Datu (leg. J.J. Vermeulen, V 1241); limestone hill on North bank Segama River, near bridge of road Sandakan to Lahad Datu (leg. J.J. Vermeulen, V 7514); Sabahmas Cave (leg. J.J. Vermeulen, V 7466, BOR/MOL 793). Tawau Hills Park (leg. J.P. King, BOR/MOL 2362). West Coast Province. Kinabalu N.P., Headquarters area near entrance (leg. J.J. Vermeulen, V 9774); Poring Hot Spring (leg. M. Schilthuizen & P. Koomen, BOR/MOL 789, BOR/MOL 790).

Description.

Shell (very) small, thin, slightly translucent or more or less opaque, pale (yellowish) brown, conical with flat or slightly convex sides; apex rounded. Surface shiny. Whorls moderately convex, the last whorl angular at the periphery, rounded above and below the periphery. Protoconch whorls convex, with fine, densely but slightly irregularly placed radial riblets; spiral sculpture about equally strong, fine, slightly more spaced grooves which are partly continuous, and partly broken up into minute pits. Teleoconch: Last whorl with or without a peripheral spiral thread coinciding with the suture of the penultimate whorl, above the periphery with numerous (rather) fine, well-spaced grooves which consist of rows of densely placed pits which are often elongated in the direction of the growth lines; below the periphery usually with numerous fine, well-spaced, continuous grooves. Radial sculpture: above the periphery with some irregularly spaced, somewhat raised growth lines, these less conspicuous below the periphery. Umbilicus open, very narrow. Dimensions: Height up to 6 mm; width up to 5.4 mm; diameters of the first four whorls 0.7-1.0 mm, 1.15-1.60 mm, 1.7-2.2 mm, 2.1-3.2 mm respectively; number of whorls up to 7 7/8, height aperture up to 2.1 mm; width aperture up to 3 mm.

Habitat in Sabah and distribution.

Primary and secondary forest on limestone and granodiorite soil, up to 1700 m alt. Sabah: Mount Kinabalu; Interior; East coast. Also in Sarawak; Kalimantan. Distribution elsewhere: Sulawesi.

Cross diagnosis.

Uniquely identified by the pitted spiral sculpture on the teleoconch, in combination with its size. In Kaliella dendrophila , of group 1, the spiral sculpture consist of beads on the shell surface, not pits in the shell surface. It is much smaller, has a thinner, more translucent shell with a smaller protoconch with predominant spiral sculpture, and with whorls more rapidly increasing in size.

Remarks.

The spiral sculpture above the periphery is fine in some specimens, somewhat coarser in others. The continuous spiral striation below the periphery is absent in some specimens.

Etymology.

Refers to the shell sculpture [punctatus (L.) = dotted].