Atholus bifrons ( Marseul, 1854 )

Cruz, Ian Niel dela, Hartini, Sri, Dwibadra, Dhian & Ôhara, Masahiro, 2024, Contribution to the Knowledge of the Genus Atholus (Coleoptera: Histeridae: Histerinae: Histerini) from the Indonesian Archipelago, Species Diversity 29 (1), pp. 119-139 : 120-121

Atholus bifrons ( Marseul, 1854)

( Figs 1A , 11D )

Hister bifrons Marseul, 1854: 545 (type locality: India).

Atholus bifrons : Lewis 1906: 402 (listed); Mazur 1984: 211; Mazur 1997: 129 ( India, Indonesia: Sumatra, Vietnam, Thailand, Hongkong); Mazur 2008: 90; Mazur 2009: 113; Mazur 2011: 103 (additional records: Taiwan; catalogued); dela Cruz and Ôhara 2022: 237–245 ( Japan: Ryukyus, Indonesia: Borneo; SEM, male and female genitalia).

Hister (Atholus) bifrons : Bickhardt 1910: 53; Bickhardt 1917: 193 (catalogued); Desbordes 1917: 323 (keyed).

Specimens examined. TwoJJ, 2♀♀, and 2 specimens of undetermined sex. Sumatra Island: Central Sumatra, Payakumbuh , Harau Valley , March to September 1993, Sarimudanas leg . (1♀ SEHU, 1♀ MZB) . Borneo Island : West Kalimantan, Mt . Bawang, 250–300 m a .s .l ., October 1990, unknown collector [IC-22-id01] (1J and 1 ex. SEHU, 1J and 1 ex. MZB) .

Diagnosis. Atholus bifrons can be recognized by the forehead divided into two by rather weak median longitudinal impression and remarkable punctations of both propygidium and pygidium.

Distribution. India, Thailand, Vietnam, China: Hong Kong, Taiwan, Ryukyu islands of Japan, and Indonesia: Sumatra and Borneo Islands ( Fig. 11D ).

Remark. The Indonesian distribution of this species previously recorded only from Sumatra has been recently extended to Borneo (dela Cruz and Ôhara 2022).

Figures

Fig. 1. Some Indonesian Atholus, dorsal habitus. A, Atholus bifrons (Marseul, 1854), IC-22-in01; B, A. philippinensis (Marseul, 1854), IC- 22-in06; C, A. coelestis (Marseul, 1857), IC-22-in02; D, A. myrmidon (Marseul, 1861), IC-22-in05; E, A. famulus (Lewis, 1892), IC-22-in03; F, A. singalanus (Marseul, 1880), IC-22-in07; G, A. torquatus (Marseul, 1854), IC-22-in08. Scale bar: 1.00mm.

Fig. 11. Map showing the occurrence of some species of Atholus from the Indonesian archipelago and its extra-limital distribution in the Oriental Region. A, Atholus coelestis; B, A. philippinesis; C, A. torquatus; D, A. bifrons.