Xoridesopus infuscatus Gupta & Gupta, 1983

Kyohei Watanabe, 2019, Taxonomic and Zoogeographical Notes of Japanese Cryptinae (Hymenoptera, Ichneumonidae), with Description of Five New Species, Bulletin of the Kanagawa Prefectural Museum (Natural Science) 48, pp. 81-113 : 108-109

Xoridesopus infuscatus Gupta & Gupta, 1983

( Figs. 13A–F ) Xoridesopus infuscatus Gupta & Gupta, 1983: 121 .

Description. The original description of this species (Gupta & Gupta, 1983) lacks some important character states. In addition, the male of this species undescribed. Thus, I redescribe this species with the character states of males below.

Female (n=5). Body length 8.5–13.5 mm, mat, covered with silver setae.

Head 0.6–0.65 times as long as wide. Clypeus 0.5 times as long as wide, slightly convex, sparsely punctate dorsally, its anterior margin with a minute median tooth. Face 0.5–0.6 times as long as wide, slightly convex medially. Frons slightly concave above antennal socket, sparsely punctate laterally, smooth ventrally, with irregular rugae mediodorsally. Occipital carina complete, its lower end joined with hypostomal carina. MSL 0.5–0.7 times as long as BWM. OOL 1.3–1.5 times as long as OD. POL 0.9 times as long as OD. Antenna with 23 flagellomeres. F1 1.05 times as long as F2.

Epomia present, weak and short. Lateral area of pronotum largely covered with longitudinal striae ( Fig. 13A ), punctate dorsally and ventrally. Notauli distinct and deep on anterior 0.6 of mesoscutum. Mesoscutum densely punctate except for a median area with irregular rugae. Scutellum slightly convex, punctate. Epicnemial carina distinct, its dorsal end close to subalar prominence. Mesopleuron with striae along epicnemial carina, subalar prominence and speculum. Speculum smooth. Metapleuron covered with irregular rugae. Propodeum with anterior transverse carina, posterior transverse carina and pleural carina, covered with irregular rugae except for area anterior to anterior transverse carina densely punctate. Propodeal apophysis absent. Propodeal spiracle oval. Fore wing length 6.5–11.0 mm. Areolet medium-sized, hexagonal, with sides convergent anteriorly. Nervulus based on basal vein. Nervellus intercepted posterior to midpoint. Fore tibia with a bulge except for basal 0.25. Hind femur 5.2–5.3 times as long as maximum depth in lateral view. Hind TS2: TS1, TS3, TS4 and TS5 = 2.0: 5.0–5.4, 1.1–1.2, 0.8–0.9 and 1.9–2.0.

Metasoma densely punctate. T1 1.65–1.85 times as long as maximum width, with a pair of small triangular teeth at base. Median dorsal carina and dorsolateral carina of T1 complete. T2 0.85–0.9 times as long as maximum width. Posterior margin of T8 truncate, its dorsal corner almost right angle in lateral view. Ovipositor sheath 0.9–1.0 times as long as hind tibia. Apex of ovipositor similar to X. amamiensis and X. tokarensis .

Coloration ( Figs. 13A–D ). Body (excluding wings and legs) black, except for: clypeus, face, mandible except for apex, all orbits (sometimes except for dorsal end of temporal orbit), gena, malar space, palpi, collar, dorsal margin of pronotum, a median spot of mesoscutum, scutellum, postscutellum, tegula, subalar prominence, speculum, a large ventral spot of mesopleuron, a large spot of mesosternum, two spots of metapleuron, a spot of propodeum, posterior band of T1–T6 and T8, lateral side of T3–T8 whitish yellow to yellow; F6–F16 (sometimes also F17 and F18) with a white band; apical part of antenna slightly tinged with reddish brown; ovipositor reddish brown. Wings hyaline; veins and pterostigma blackish brown to brown except for yellow wing base. Fore and mid legs whitish yellow to reddish yellow except for TS4 and TS5 tinged with blackish brown. Hind leg whitish yellow, except for: coxa largely black, trochanter and trochantellus tinged with blackish brown; hind femur reddish brown except for brown apex; hind tibia reddish brown to blackish brown; TS4 and TS5 blackish brown.

Male (n=3). Similar to female ( Figs. 13E, F ). Body length 9.0 mm. Face 0.6–0.7 times as long as wide. OOL 1.2–1.3 times as long as OD. POL 0.8–0.9 times as long as OD. Antenna with 26 flagellomeres. F1 1.05–1.1 times as long as F2. F13 and F14 (sometimes also F15) with a tyloid. Fore wing length 6.5 mm. Fore tibia without a bulge. Hind femur 5.4–5.8 times as long as maximum depth in lateral view. Hind TS2: TS1, TS3, TS4 and TS5 = 2.0: 3.4–3.6, 1.3–1.5, 0.7–0.8 and 0.8–0.9. T1 2.85–3.0 times as long as maximum width. T2 2.0–2.15 times as long as maximum width. Antenna without a white band. Yellow area of mesopleuron and mesosternum partly united into a single large area. Base of T1 narrowly yellow to whitish yellow. Posterior margin of T1–T6 with a white band.

Specimens examined. JAPAN: AEIC, F (holotype), Okinawa Pref., Okinawa Is., Chizuka, July-September, Bohart & Harnage leg. ; KPM-NK 5006288 , F, Okinawa Pref., Okinawajima Is., Nago City, Mt. Nagodake , 13. V. 2002, H. Irei & H. Makihara leg. (MsT) ; KPM-NK 5006284 , 5006289 , 1 F & 1 M, ditto, 12–16. IX. 2004 ; KPM-NK 5006290 , M, ditto, 18. IX. 2002 ; KPM-NK 5006286 , M, Okinawa Pref., Kunigami Vil. , Uka, 16. VIII. 2001, H. Irei & H. Makihara leg. (MsT) ; KPM-NK 5006287 , F, Okinawa Pref., Kunigami Vil. , Ginama, 16. VIII. 2001, H. Irei & H. Makihara leg. (MsT) ; KPM-NK 5006283 , F, Okinawa Pref., Kunigami Vil. , Oku, 16. VIII. 2001, H. Irei & H. Makihara leg. (MsT).

Distribution. Japan (Okinawajima Is.).

Remarks. This is the first record of the male of this species.

Figures

Fig. 13. Xoridesopus infuscatus Gupta & Gupta, 1983 (A–F) and X. tokarensis sp. nov. (G–K), KPM-NK 5006283 (A–D), 5006289 (E, F) and 69548 (G–K), females (A–D, G–K) and male (E, F) ― A, F, habitus, lateral view; B, G, head, mesosoma and metasoma, dorsolateral view; C, H, head, frontal view; D, head and mesosoma, lateral view; E, nervellus of hind wing.