Riekoleon, New, 1985

Machado, Renato Jose Pires & Oswald, John David, 2020, Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae), Zootaxa 4796 (1), pp. 1-322 : 286-287

Key to species of Riekoleon

1 Profemur ≥ 3.5x length of procoxa; male ectoproct with a tubularly-produced posteroventral lobe ( Fig. 152a )....................................................................................... ( convergens species group) … 6

- Profemur <3.5x length of procoxa; male ectoproct without a tubularly-produced posteroventral lobe ( Fig. 166a )..........................................................................................( rudda species group) … 2

2 (1’) Forewing: first fork of RP in proximal half of wing ( Fig. 165a ); female anterior gonapophysis present (a small rounded plate) ( Figs. 166 d–e)........................................................................................ 3

- Forewing: first fork of RP in distal half of wing ( Fig. 154c ); female anterior gonapophysis absent ( Figs. 155 d–e)............................................................................................. R. distivenus sp. nov.

3 (2) Ocular setae absent; pronotum as long (or longer) than wide; male mediuncus absent ( Figs. 166 b–c)................... 4

- Ocular setae present (white); pronotum wider than long ( Fig. 169b ); male mediuncus present (but small) ( Fig. 170b )................................................................................................... R. striatus

4 (3) Forewing: dark mediocubital line continuing as a long dark streak across radial area; hind wing rhegmal area with a large brown spot ( Fig. 160c )....................................................................................... 5

- Forewing: dark mediocubital line not continuing as a long dark streak across radial area; hind wing rhegmal area without a large brown spot (only a small dark infuscation) ( Fig. 165c ).................................................. R. rudda

5 (4) Hind wing with apical spot as large as rhegmal spot ( Fig. 160c ); pronotum mostly pale, but with a dark sagittal stripe ( Fig. 160b ); profemur mostly pale; female posterior gonapophysis longer than wide ( Figs. 161 a–b)...... R. longitudinalis sp. nov.

- Hind wing with apical spot much smaller than rhegmal spot ( Fig. 156c ); pronotum mostly dark ( Fig. 156b ); profemur entirely black; female posterior gonapophysis wider than long ( Fig. 157f )....................................... R. edwardsi

6 (1) Pterothorax and abdomen without scale-like setae........................................................... 7

- Pterothorax and abdomen with scale-like setae ( Fig. 167b ).................................... R. squamosus sp. nov.

7 (6) Tibial spurs present.................................................................................... 8

- Tibial spurs absent............................................................................. R. proctus

8 (7) Body dark (vertex, mesonotum and legs mostly brown) ( Fig. 158b ); pronotum setae mostly black and short; hind wing with rhegmal spot large ( Fig. 158c )................................................................... R. furcatus

- Body pale (vertex, mesonotum and legs mostly pale) ( Fig. 151b ); pronotum setae mostly white and long; hind wing with rhegmal spot small ( Fig. 151c ).................................................................... R. convergens

Figures

FIGURE 151. Riekoleon convergens: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal (ends of legs showing through wings at base).

FIGURE 152. Riekoleon convergens: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female: d) terminalia, lateral; e) terminalia, posteroventral (note pregenital plate hidden beneath sternite). Scale bars = 0.5 mm.

FIGURE 154. Riekoleon distivenus sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 155. Riekoleon distivenus sp. nov.: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female: d) terminalia, lateral; e) terminalia, ventral. Scale bars = 0.5 mm.

FIGURE 156. Riekoleon edwardsi: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 157. Riekoleon edwardsi: male: a) terminalia, lateral; b) 9th sternite, ventral; c) genitalia, lateral; d) genitalia, posterior; female: e) terminalia, lateral; f) terminalia, ventral (note pregenital plate hidden beneath sternite); g) pregenital plate, posterior. Scale bars = 0.5 mm.

FIGURE 158. Riekoleon furcatus: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 161. Riekoleon longitudinalis sp. nov.: female: a) terminalia, lateral; b) terminalia, ventral (note pregenital plate hidden beneath sternite). Scale bars = 1mm.

FIGURE 165. Riekoleon rudda: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 166. Riekoleon rudda: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female: d) terminalia, ventral (note pregenital plate hidden beneath sternite); e) terminalia, lateral. Scale bars = 0.5 mm.

FIGURE 167. Riekoleon squamosus sp. nov.: a) head, anterior; b) abdomen base, lateral (note scale-like setae); c) head and thorax, dorsal; d) wings, dorsal.

FIGURE 169. Riekoleon striatus: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 170. Riekoleon striatus: male: a) terminalia, lateral; b) genitalia, posterior; c) genitalia, lateral; female: d) terminalia, lateral; e) terminalia, ventral. Scale bars = 0.5 mm.