Heterometrus laoticus Couzijn, 1981

Prendini, Lorenzo & Loria, Stephanie F., 2020, Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005, Bulletin of the American Museum of Natural History 2020 (442), pp. 1-480 : 245

Heterometrus laoticus Couzijn, 1981

Figures 7F, 8F, 10 , 24A, B , 38A, B , 51A–D , 70A , 71A , 72A , 157 , 169–173, table 2

Palamnaeus silenus Simon, 1884a: 361 (misidentification, part); Thorell, 1889: 588 (misidentification, part); Flower, 1901: 35, 36 (misidentification); Simon, 1904: 293 (misidentification, part).

Heterometrus (Scorpio) longimanus: Kraepelin, 1901: 271 (misidentification, part).

Heterometrus (Scorpio) longimanus var. silenus:

Figures

FIGURE 10. Phylogeny of Asian forest scorpions (Scorpionidae: Heterometrinae Simon, 1879, stat. nov.), obtained by simultaneous phylogenetic analysis of 186 morphological characters and 4188 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci (Loria and Prendini, in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 1).

FIGURE 24. Heterometrus Ehrenberg, 1828, carapace, dorsal aspect. A, B. Heterometrus laoticus Couzijn, 1981, A. ♂ (AMNH), Phahom Camp, Laos, B. ♀ (AMNH), Tham Pha In, Laos. C, D. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev.C. ♂, D. ♀ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 2 mm.

FIGURE 38. Heterometrus Ehrenberg, 1828, sternum and pectines, ventral aspect. A, B. Heterometrus laoticus Couzijn, 1981, A. ♂ (AMNH), Phahom Camp, Laos, B. ♀ (AMNH), Tham Pha In, Laos. C, D. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev. C. ♂, D. ♀ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 2 mm.

FIGURE 51. Heterometrus Ehrenberg, 1828, legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. E–H. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. Scale bars = 2 mm.

FIGURE 70. A–D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev., metasoma and telson, dorsal aspect. A. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. B. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. D. Heterometrus thorellii (Pocock, 1892), ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. E. Javanimetrus cyaneus (Simon, 1872), comb. nov., ♂ (AMNH), Ujung Kulon, Indonesia. Scale bars = 5 mm.

FIGURE 71. A–D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev., metasoma and telson, lateral aspect. A. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. B. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. D. Heterometrus thorellii (Pocock, 1892), ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. E. Javanimetrus cyaneus (Simon, 1872), comb. nov., ♂ (AMNH), Ujung Kulon, Indonesia. Scale bars = 5 mm.

FIGURE 72. A–D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev., metasoma and telson, ventral aspect. A. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. B. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. D. Heterometrus thorellii (Pocock, 1892), ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. E. Javanimetrus cyaneus (Simon, 1872), comb. nov., ♂ (AMNH), Ujung Kulon, Indonesia. Scale bars = 5 mm.

FIGURE 157. Map of Southeast Asia, plotting known locality records of three species of Heterometrus Ehrenberg, 1828, based on material examined, and verified records from virtual museums and the literature: Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev.; Heterometrus laoticus Couzijn, 1981; Heterometrus spinifer (Ehrenberg, 1828).