Orphninae

Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A., 2016, Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood, Zootaxa 4207 (1), pp. 1-93 : 12

Key to the Madagascar Orphninae genera (both sexes)

1. Protibiae with 2 outer teeth apically and 1 much smaller outer tooth basally ( Fig. 7 D)........... Pseudorphnus Benderitter

- Protibiae with 3 outer teeth in apical half ( Figs. 10 H, 30C, 28F).................................................2

2. Body length smaller (usually less than 7 mm); lateral margin of pronotum with 4 long setae: 2 near anterior margin, 1 in the middle, and 1 near posterior margin; some or all these setae may be abraded, but 4 chaetiferous punctures are always distinct; left and right mandibles may differ strongly in length in males, and frontoclypeus is somewhat asymmetrical in such cases; disc of pronotum smooth, without processes or excavations.................................. Madecorphnus Paulian

- Body length larger (usually longer than 7 mm); lateral margin of pronotum with numerous setae (at least more than 10) evenly distributed along the margin, setae may differ strongly in length; left and right mandibles of about same length, frontoclypeus symmetrical; disc of pronotum may be with more-or-less developed tubercles, ridges, and excavations in males........... 3

3. Eyes smaller (width of eye in dorsal view more than 7 times smaller than distance between eyes); propleurae with a longitudinal ridge bearing a row of setae parallel to lateral margin of pronotum; disc of pronotum with more-or-less developed tubercles, ridges, and excavations in males, and smooth in females; apical outer tooth of protibia in males directed laterally, at reasonable angle to internal margin of tibia ( Fig. 30 C)....................................... Triodontus Westwood

- Eyes larger (width of eye in dorsal view less than 4 times smaller than distance between eyes); propleurae without a longitudinal ridge, only with a sparser row of setae; disc of pronotum with 2 feebly developed tubercles in middle or without tubercles; apical outer tooth of protibia almost parallel to internal margin of tibia ( Fig. 28 F)........ .. Renorphnus Frolov & Montreuil

Figures

FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H).

FIGURE 10. Madecorphnus montreuili. Habitus in dorsal view (A—male, C—female), aedeagus in lateral view and parameres in dorsal view (B), labels of holotype (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G), protibia (H, male left, female right).

FIGURE 28. Renorphnus clementi. Habitus in dorsal view (A—male, B—female), aedeagus in lateral view and parameres in dorsal view (C), internal sac of aedeagus (D), distributional records (E), protibia (F, male left, female right).

FIGURE 30. Triodontus hova. Abdomen of male (A), syntype of O. nigrita, habitus in dorsal view (B), protibia (C, male left, female right).