Ummidia erema (Chamberlin, 1925)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia erema (Chamberlin, 1925) Figs 64 , 65 , Map 5

Pachylomerus eremus Chamberlin, 1925d: 211; HOLOTYPE: 1 ♀ (IZ16003) from Barro Colorado Island, Panama Canal Zone, Panama, 9.1520 -79.8467 5, 156 m a.s.l., coll. WC Allee, deposited in the MCZ, examined.

Diagnosis.

Ummidia erema are relatively small spiders (CL 3.8-4.6) with distinctly light tarsi. Males can be distinguished from U. zilchi , U. carlosviquezi , U. varablanca , and U. quepoa by possessing a sinuous embolus, rather than one with an even single curve and from U. yojoa and U. varablanca by the presence of many prolateral (15 vs 0) and retrolateral (21 vs 2 or 3) spines on tibia I. Males can be further differentiated from U. rugosa and U. riverai by the lack of a pale dorsal heart patch on the abdomen. Females can be distinguished from all geographically proximate species except U. hondurena and U. rugosa by having spermathecae which bend first medially and then anteriorly.

Description of female holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4, leg IV tarsi brownish yellow 10YR 6/8. Abdomen dark brown 2.5YR 3/2, spinnerets yellow 10YR 7/6. Cephalothorax. Carapace 3.79 long, 3.68 wide. Pars cephalica 2.83 long. Foveal groove procurved, 0.82 long, 0.3 wide. Eye tubercle moderate. AER procurved. PER relatively straight. Eye group 1 long, 0.58 wide, AME 0.22, PME 0.2, ALE 0.34, PLE 0.28. Sternum sparsely setose, thicker around anterior edges, STRl 2.32, STRw 2.25. Chelicerae with anterior row comprising three teeth, posterior margin with six teeth. Palpal endites with 27 large cuspules across proximal half of endite face and eleven cuspules distally, ENDw 0.95, ENDl 1.66. Labium with eight cuspules; five large and three small, LBw 0.9, LBl 0.63. Rastellum with six spines along distal cheliceral margin and five spines medially on process. Abdomen. Evenly setose. Legs. F1 2.88; F1w 0.99; P1 1.71; Ti1 1.66; Mt1 1.19; Tr1 0.82; F3 2.31; F3w 1.12; P3 1.33; Ti3 1.26, Sd3 0.87; Mt3 1; Tr3 0.98; F4 2.86; F4w 1.15; P4 1.46; Ti4 1.54; Mt4 1.19; Tr4 0.82. Retrolateral face tarsus IV with defined comb of alternating long and short spinules. Pedipalps. PF 2.66, PP 1.43, PTi 1.43, PTr 1.57. Spermathecae bent medially and then anteriorly, bulbs facing anteriorly.

Variation, females

(n = 3). CL 3.79-4.59, 4.16 ± 0.23; CW 3.68-4.38, 3.95 ± 0.21; Cap 2.83-3.31, 3.06 ± 0.14; ENDl 0.58-0.65, 0.61 ± 0.02; ENDw 0.98-1.06, 1.01 ± 0.02; STRl 2.26-2.6, 2.39 ± 0.11; STRw 2.25-2.67, 2.4 ± 0.14; LBl 0.63-0.81, 0.74 ± 0.06; LBw 0.9-1.05, 0.96 ± 0.04; F1 2.51-3.11, 2.83 ± 0.18; F1w 0.94-1.1, 1.01 ± 0.05; P1 1.34-1.82, 1.62 ± 0.15; Ti1 1.37-1.86, 1.63 ± 0.14; Mt1 1.06-1.29, 1.18 ± 0.07; Tr1 0.72-0.87, 0.8 ± 0.04; F3 2.26-2.65, 2.4 ± 0.12; F3w 1.12-1.27, 1.18 ± 0.05; P3 1.18-1.5, 1.34 ± 0.09; Ti3 1.25-1.38, 1.3 ± 0.04; Mt3 0.92-1.12, 1.01 ± 0.06; Tr3 0.98-1.09, 1.04 ± 0.03; F4 2.78-3.24, 2.96 ± 0.14; F4w 1.11-1.34, 1.2 ± 0.07; P4 1.41-1.78, 1.55 ± 0.12; Ti4 1.34-1.69, 1.52 ± 0.1; Mt4 1.31-1.58, 1.45 ± 0.08; Tr4 0.9-1.02, 0.95 ± 0.04; PF 2.53-2.94, 2.71 ± 0.12; PP 1.26-1.62, 1.44 ± 0.1; PTi 1.42-1.58, 1.49 ± 0.05; PTr 1.45-1.65, 1.56 ± 0.06.

Description of male exemplar.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp and leg I removed, in vial with specimen. General coloration. Carapace, chelicerae, and legs reddish black 10R 2.5/1, tarsi reddish yellow 7.5YR 6/8. Abdomen black 5YR 2.5/1. Cephalothorax. Carapace 3.9 long, 3.83 wide. Pars cephalica 2.75 long. Foveal groove procurved, 0.31 long, 0.64 wide. Tubercle high. AER procurved. PER straight. Eye group 0.59 long, 1.03 wide, AME 0.35, PME 0.19, ALE 0.37, PLE 0.28. Sternum sparsely setose around outer 1/3, STRl 2.19, STRw 2.1. Chelicerae with anterior tooth row comprising five teeth, posterior margin with seven teeth. Palpal endites with 28 hastate cuspules of varying size across proximal 3/4 of endite face, lacking distal endite cuspules, ENDw 0.9, ENDl 1.5. Labium with ten cuspules, LBw 0.83, LBl 0.55. Rastellum greatly reduced with six small cuspules. Abdomen setose. Legs. F1 3.88; F1w 0.87; P1 1.88; Ti1 2.74; Mt1 1.83; Tr1 0.98; F3 2.95; F3w 1.07; P3 1.41; Ti3 1.72; Sd3 1.11; Mt3 1.46; Tr3 1.13; F4 3.74; F4w 0.98; P4 1.64; Ti4 2.19; Mt4 2.18; Tr4 1.23. Retrolateral face of tarsus IV lacking defined brush or comb. Leg I spination pattern: TSp 8, TSpv 7, TSrd 0, TSr 1, TSrv 20, MtSp 7, MtSr 13, TrSp 4, TrSr 7. Pedipalps. PTl 2.12, PTw 0.78, Bl 1.87. Embolus relatively long, slightly sinuous.

Variation, males.

Known only from male exemplar.

Material examined.

Panama: Panama Canal Zone : Barro Colorado Island, 9.152 -79.8467 5, 156 m a.s.l. (MCZ1261, 1♀, WC Allee, MCZ) (UMM0514, 1946, 1♀, Zetek, AMNH) (UMM0512, 1.i.1947, 1♀, NLH Krauss, AMNH); (UMM0535, 18.iv.1946, 1♂, TC Schneirla, AMNH); Ft. San Lorenzo, 9.3227 -80.0014 4, 15 m a.s.l. (UMM0513, 1juv, Zetek, AMNH) .

Figures

Map 5. Locality records for southern Mexico and Central American species.

Figure 64. Ummidia erema (Chamberlin, 1925) from Barro Colorado Island, Panama. A male habitus illustration UMM 535 B female habitus illustration MCZ 16003. Scale bars: 4.0 mm.

Figure 65. Ummidia erema (Chamberlin, 1925) from Barro Colorado Island, Panama. A-D male exemplar (UMM 535) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female holotype (MCZ 16003). Scale bars: 1.0 mm.