Mastigoproctus Pocock, 1894

Barrales-Alcalá, Diego, 2018, Systematic Revision Of The Giant Vinegaroons Of The Mastigoproctus Giganteus Complex (Thelyphonida: Thelyphonidae) Of North America, Bulletin of the American Museum of Natural History 2018 (418), pp. 1-64 : 8-9

Mastigoproctus Pocock, 1894

Figures 1 –20, tables 1–5

Mastigoproctus Pocock, 1894: 129 , 130; Kraepelin, 1897: 36; Kraepelin, 1899: 223; Pocock, 1902a: 46; Shipley, 1909: 312; Hirst, 1912: 237; Hartline, 1923: 148, 149; Mello-Leitão, 1931: 25; Werner, 1935: 468; Takashima, 1947: 42; Snodgrass, 1948; 12, 27; Roewer, 1954: 57; Besch, 1969: 728; Weygoldt, 1972a: 23, 24, 29, 44, 45, 48; Rowland and Cooke, 1973: 68; Phillips, 1976: 397, 401, 403; Weygoldt, 1978: 145, 146, 154, 155; Valerio, 1981: 15; Mann, 1984: 149, 150; Legendre, 1985: 44; Haupt et. al., 1988: 883; Weygoldt, 1988: 189, 190, 194; Ruppert and Barnes, 1994: 334; Barriel and Tassy, 1998: 199, fig. 3; Dunlop, 1998: 291, 293, 294; Proctor, 1998: 160; Armas and Maes, 1999: 14; Shultz, 1999: 92, 96, 104, 105, 109, 111; Armas, 2000: 2; Ax, 2000: 103, 390; Adis et al., 2002: 8; Armas, 2002: 42; Dunlop and Martill, 2002: 329; Harvey, 2002: 363; Rowland, 2002: 194, 195; Giribet, 2003: 558, fig. 1; Harvey, 2003: 65; Armas, 2004: 23, 24; Regier et al., 2005: 396, fig. 1; Víquez and Armas, 2005: 95, 96, 98; 2006: 37; Ballesteros and Francke, 2006: 156; Punzo, 2006: 266; Víquez and Armas, 2007: 39–42; Armas and Víquez, 2007: 508; Dunlop et al., 2007: 124, 125; Huff et al., 2008: 1, 2; Dunlop and Tetlie, 2008: 551; Giupponi and De Vasconcelos, 2008: 18; Tetlie and Dunlop, 2008: 299; Teruel and Armas: 2008: 32; Armas et al., 2009: 4; Beccaloni, 2009: 113, 115–117, 124; Haupt, 2009: 15, 18; Villareal and Giupponi, 2009: 145, 146; Lukhtanov and Kuznetsova, 2010: 1117, fig. 2; Teruel, 2010: 193; McMonigle, 2013: 7, 10, 12, 16, 26, 28, 30, 31, 49–52, 55–57, 59, 65, 68, 77, 80–82, 85, 88, 89, 91, 94, 98, 99, 102, 106, 107, 112, figs. unnumbered, 4–6, 8, 9, 14, 27–29, 33, 34, 36, 37, 47, 49, 53, 54, 58, 60, 61, 66, 68, 79, 82–84, 86, 90, 91, 94, 97; Weygoldt and Huber, 2013: 348: Maquart et al., 2016: 79; Monjaraz-Ruedas et al., 2016: 120; Selden et al., 2016: 7; Chenyang and Huang, 2017: 101, 104.

DIAGNOSIS: Mastigoproctus differs from Mayacentrum Víquez and Armas, 2006 , Ravilops Víquez and Armas, 2005 , and Thelyphonellus Pocock, 1894 , in the presence of a well-developed carina that extends from the lateral ocelli approximately two-thirds the length of the carapace toward the median ocular tubercle. Mastigoproctus differs further from Thelyphonellus in the presence of a pair of pygidial ommatoids, which are absent in the latter genera, and from Ravilops by the shape and macrosculpture of the pedipalps, which are elongate, punctate, and tuberculate in Mastigoproctus , but short and smooth in Ravilops . Additionally, Mastigoproctus differs from Mayacentrum in the structure of the tergites and sternites. Only the anterior tergites (I–III) are divided medially in Mastigoproctus , whereas all tergites are partially or completely divided in Mayacentrum , and sternites II and III are unmodified or with at most vague lateral swellings in the adult male of Mastigoproctus , whereas sternites II and III are distinctly bilobed in the adult male of Mayacentrum .

Mastigoproctus differs from Mimoscorpius Pocock, 1894 , as follows. The pedipalp femur of the adult male bears a pair of teeth, one on the retroventral surface and one on the proventral surface, in Mastigoproctus whereas only one tooth is present on the proventral surface in Mimoscorpius . The pedipalp tibia of the adult male is distinctly longer than wide in Mastigoproctus but almost as wide as long in Mimoscorpius . The tibiae of legs II–IV each possess a single ventrodistal spur in Mastigoproctus whereas two ventrodistal spurs are present in Mimoscorpius . The seminal receptacles are short, triangular, and posteriorly directed in Mastigoproctus but tubular and kidney shaped in Mimoscorpius .

Mastigoproctus differs from Valeriophonus Víquez and Armas, 2005 , as follows. The tibiae of legs II–IV each possess a single ventrodistal spur in Mastigoproctus whereas a ventrodistal spur is present only on the tibia of leg IV in Valeriophonus . The pedipalp of Mastigoproctus is elongate and without a lobe on the retrolateral surface of the fixed finger, whereas the pedipalp of Valeriophonus is short and robust, with a pronounced lobe on the retrolateral surface of the fixed finger. The macrosetae comprising the stridulatory surface of the pedipalp coxa are randomly scattered in Mastigoproctus but evenly aligned in Valeriophonus .