Apterodela (s. str.) kazantsevi ( Matalin, 2001 )

Matalin, Andrey V., Wiesner, Jürgen, Xiong, Xinxin & Araki, Takashi, 2024, Revision of the genus Apterodela Rivalier, 1950 (Coleoptera, Cicindelidae), Zootaxa 5405 (3), pp. 301-353 : 318-322

Apterodela (s. str.) kazantsevi ( Matalin, 2001)

Figs 25, 26 , 54, 55 , 93, 98 , 133, 138 , 149 , 201, 211 , 222 , 233 , 239 , 246 , 253 , 260 .

Cylindera (Apterodela) kazantsevi Matalin, 2001: 385 (Type locality— China, Jiangxi, Jinggang Shan ).

Cicindela lobipennis Bates, 1888 : Fang & Wu 2001: 70.

Cylindera (Apterodela) kazantsevi Matalin, 2001 : Lorenz 2005: 165; Shook & Wiesner 2006: 13; Wu & Shook 2010: 81; Wu 2011: 26.

Apterodela kazantsevi ( Matalin, 2001) : Putchkov & Matalin 2017: 217; Wiesner 2020: 266.

Type material. Holotype, ♂ ( Figs 25, 26 )— China, Jiangxi, Jinggangshan , h = 700–1000 m, 25–31.V.1998, leg. S. Kazantsev [white printed label] , HOLOTYPE, Cylindera (Apterodela) kazantsevi sp. nov., det. A.V. Matalin, 2001 [red printed label] (cSK) ; PARATYPE, 1♂ — ibid , PARATYPE, Cylindera (Apterodela) kazantsevi sp. nov., det. A.V. Matalin, 2001 [red printed label] (cSK) .

Addition material. China, Zhejiang Province: 1 ♂, 1 ♀ — China, Zhejiang, Baima Shan , 7– 17.06.2008 (cTA) ; 1 ♂ — ibid (cMH) ; 1 ♂ — ibid (cJM) ; 1 ♂, 2 ♀♀ — China, SW Zhejiang, Fengyangshan, Huangmao Jian, 27 o 53’N 119 o 11’E, 12.V–5.VI.2008, leg. Jaroslav Turna ; 1 ♂ — ibid, 4.V–2.VI.2009, leg. Jaroslav Turna ; 1 ♀ — China, Zhejiang, Yunhe, h = 500 m, Lishui , VI.2021 (all TA) ; 4 ♂♂, 4 ♀♀ (digital images)— China, SW Zhejiang, Fengyangshan, Huangmao Jian, 27 o 53’N 119 o 11’E, 12.V–5.VI.2008, leg. Jaroslav Turna (cZS) ; Jiangxi Province: 1 ♂, 1 ♀ — China, NE Jiangxi, Wufugang, 1400–1500 m, E slopes, 27 o 59’N 118 o 02’E, 29.IV–27.V.2010, leg. Jaroslav Turna (cJG) ; Fujian Province: 1 ♂, 1 ♀ — China, centr. Fujian, Tianbaoyanshan, NW slopes, 1100–1200 m, 25°58’N 117°32’E, 21.V–26.VI.2011, leg. Jaroslav Turna (cJW) ; Guangdong Province: 3 ♀♀ — China, NE Guangdong prov., Quifeng Shan, 900–1200 m, 23°51’N 115°21’E, 31.V–18.VI.2015, leg. J. Turna (cJW) .

References. China, Zhejiang Province: 1 ♀ (digital image)— China, Zhejiang, Quzhou , 28.786191N 119.064983E, 17.IV.2021, Amaël Borzée (https://guatemala.inaturalist.org/observations/74219090); 1 ♀ (digital image)— China, Zhejiang, Quzhou , 28.786198N 119.064957E, 17.IV.2021, Yoonjung Yi (https://guatemala. inaturalist.org/observations/74228298); 1 ♂ (digital image)— China, Zhejiang, Lishui, Longquan Mt , 28.070828N 118.80722E, 11.VII.2023 (https://guatemala.inaturalist.org/observations/172232916); Fujian Province: 1 ♂ (digital image)— China, Fujian, Nanping, Wuyishan Mt , 27.755707N 117.74285E, 19.VII.2020 (https://guatemala. inaturalist.org/observations/62631135).

Differential diagnosis. Apterodela kazantsevi is easily distinguished from A. bivirgulata by the deep subapical notch ( Figs 201, 211 vs. Figs 155–162, 165––179, 183–189 ) as well as by the shape of aedeagus ( Figs 222 , 233 vs. Figs 214–219 , 228–231 ) and internal sac ( Figs 239 , 246 , 253 , 260 vs. Figs 238 , 245 , 252 , 259 ). Apterodela kazantsevi is readily distinguished from related A. latissima sp. nov. (only by males) by the longer labrum LW/LL = 1.67–2.0 (mean = 1.89) vs. 1.97–2.38 (mean = 2.17), the narrower elytra EW/EHW = 1.44–1.69 (mean = 1.58) vs. 1.71–1.78 (mean = 1.75) with less sinuate lateral margins and longer subapical notch ( Fig. 201 vs. Figs 199, 200 ), the less convex body TL/BH = 4.15–4.59 (mean = 4.31) vs. 4.0–4.28 (mean = 4.14), and the shape of aedeagus ( Fig. 222 vs. Fig. 223 ). Apterodela kazantsevi is clearly separated from A. lobipennis by the longer labrum, especially in females LW/LL = 1.67–2.0 (mean = 1.89) vs. 1.82–2.40 (mean = 2.05) in males and 1.22–1.53 (mean = 1.31) vs. 1.50 –2.00 (mean = 1.73) in females, the narrower pronotum HW/PW = 1.39–1.6 (mean = 1.51) vs. 1.3–1.52 (mean = 1.4), the wider elytra with longer and deeper subapical notch ( Figs 201, 211 vs. Figs 193–198, 203–210 ), by the shape of aedeagus ( Figs 222 , 233 vs. Figs 220, 221 , 232 ) and internal sack ( Figs 239 , 246 , 253 , 260 vs. Figs 240 , 247 , 254 , 261 ). Apterodela kazantsevi is differentiated from A. alopecomma sp. nov. by the smaller size TL = 14.7–15.6 (mean = 15.2) mm vs. 15.5–16.7 (mean = 15.98) mm, flatter body TL/BH = 4.15–4.59 (mean =4.31) vs. 3.93–4.19 (mean = 4.04), wider head HW/PW = 1.39–1.6 (mean = 1.51) vs. 1.24–1.33 (mean = 1.29), glabrous scape ( Figs 54, 55 vs. Figs 57, 58 ), shape of labrum ( Figs 93, 98 vs. Figs 95, 100 ), wider elytra EL/EW = 1.57–1.74 (mean = 1.66) vs. 1.72–1.78 (mean = 1.75), and the shape of aedeagus ( Figs 222 , 233 vs. Figs 224 , 234 ) and internal sac ( Figs 239 , 246 , 253 , 260 vs. Figs 241 , 248 , 255 , 262 ).

Redescription. TL = 14.7–15.6 mm (mean = 15.2 mm, n = 4) in males, 15.2–18.1 mm (mean = 16.8 mm, n = 5) in females ( Fig. 25 ); body flat, slightly convex, in males weaker than in females—TL/BH = 4.15–4.59 (mean = 4.31, n = 4) vs. 3.53–3.84 (mean = 3.75, n = 5) ( Fig. 276 ; Table 1 ).

Head metallic bronze with light bluish-green and golden-cupreous lustre on clypeus, frons and anterior edge of genae, and bright purple-cupreous reflection along sides of orbital plates; glabrous, coarsely wrinkled, with thin dense frontal furrows; orbital plates with 10–11 deep furrows; vertex widely concave, in anterior portion with wide U–shaped depression especially in females; HW/PW = 1.39–1.6 (mean = 1.51, n = 10) ( Figs 274, 275, 278 ). Antennae projected posteriorly over basal third of elytra; antennomeres 1–4 metallic greenish-bronze, bronze or cupreous with golden-cupreous or greenish-gold tinge; scape besides single apical seta glabrous; antennomere 3 with three–five while antennomere 4 with two–four stout white setae except apical ones ( Figs 54, 55 ). Labrum transverse, LW/LL = 1.67–2.0 (mean = 1.89, n = 5) in males, 1.22–1.53 (mean = 1.31, n = 5) in females, unidentate with long, stout apical tooth in females and short tooth in males and 4–9 submarginal setae ( Figs 93, 98 ).

Pronotum metallic purple-bronze with golden or golden-cupreous tinge; from slightly longitudinal to quadrate, PW/PL = 0.81–1.0 (mean = 0.91, n = 12); wrinkled and practically flat on disc; with sparse irregular rows of white decumbent hairs along straight, converging to the base lateral margins; anterior sutures indistinct, posterior ones shallow; midline thin, metallic green with distinct golden-cupreous reflection and deep small pit basally ( Figs 133, 138 ). Thoracic segments deep bronze, glabrous, coarsely rugose ( Fig. 149 ). Legs metallic green with bronze-cupreous or purple-cupreous reflections.

Elytra purple-bronze with cupreous tinge, with numerous, dense diffused, small violet-green or bluish-green punctures and with humeral, subsutural, and subapical rows of large golden-green setigerous pores; flattened, EL/ EW = 1.57–1.74 (mean = 1.66, n = 12); with wide and deep subapical sinuate notch in both sexes. Shoulders narrow, sloping, EW/EHW = 1.44–1.69 (mean = 1.58, n = 7) in males, 1.66–1.86 (mean = 1.78, n = 5) in females; scutellum bronze, large; humeral group of series umbilicata consists of 5–7 large setigerous pores ( Fig. 149 ); epipleura bronze, broad. White elytral pattern presented by the small, oval sub-humeral spot and short, comma-shaped, indistinctly oblique apical portion of middle band, supra-apical spot in males entirely absent, in females virtually invisible small, narrow, stripe-shaped ( Figs 201, 211 ).

Abdominal ventrites glabrous, black with a light bluish or violet tinge along anterior margin.

Aedeagus medium-sized, EL/AL = 2.14–2.26 (mean = 2.2, n = 2), with short, slightly broadened apex terminating in indistinct knob and extended lateral flanks ( Figs 222 , 233 ). Internal sac with long straight mt, large ag, ll, and sh-ar, distinct guide for f, which placed across DA and VLL, large protruding VA and only marked both BLL and BLR ( Figs 239 , 246 , 253 , 260 ).

Distribution. CHINA: Zhejiang, Jiangxi, Fujian, Guangdong Provinces ( Fig. 280 ).

Figures

FIGURE 280. Distribution of Chinesespecies of Apterodela (red open circles—A. (Protoapterodela) shirakii; red solid circles— A. (s. str.) bivirgulata bivirgulata; red half-solid circles—A. (s. str.) bivirgulata occidentalis ssp. n.; yellow quadrates—A. (s. str.) lobipennis; light blue rhombs—A. (s. str.) kazantsevi; orange triangles—A. (s. str.) alopecomma sp. n.; lilac rectangular—A. (s. str.) latissima sp. n.; T in the symbols—type localities; L in the symbols—according only to the literature sources). Maps source for China—https://upload.wikimedia.org/wikipedia/commons/thumb/a/ad/China_LCC_topographic_map.svg/2378px- China_LCC_topographic_map.svg.png, for Taiwan—https://upload.wikimedia.org/wikipedia/commons/b/b0/Taiwan_relief_ location_map.jpg.

FIGURES 25–34. Apterodela spp., type specimens: 25, 26—A. (s. str.) kazantsevi; 27—A. (s. str.) latissima sp. nov.; 28—A. (s. str.) alopecomma sp. nov.; 29–32—A. (Protoapterodela) shirakii; 33, 34—A. (Protoapterodela) unipunctata; 25, 27–29, 31, 33—habitus; 26, 30, 32, 34—labels; 25–30, 33, 34—males; 31, 32—females; 25–28, 33, 34—holotypes; 29, 30—lectotype; 31, 32—paralectotype.

FIGURES 49–62. Apterodela spp., head: 49, 50—A. (s. str.) ovipennis (49—Japan, Honsu, Mooka; 50—Japan, Itoigawa city, Agero); 51–53—A. (s. str.) lobipennis (51—China, Jiangxi, Kiu Kiang; 52—China, Hubei, Chang Yang; 53—China, Hubei, Dabie Shan); 54, 55—A. (s. str.) kazantsevi (54—China, Jiangxi, Jinggangshan; 55—China, Jiangxi, Wufugang); 56—A. (s. str.) latissima sp. nov. (China, Yunnan, Xishangbana); 57, 58—A. (s. str.) alopecomma sp. nov. (57—China, Sichuan, Qingcheng Hou Shan; 58—China, Sichuan, Guan Xian); 59, 60—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 61, 62— A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P.); 49, 52, 54, 56, 57, 59, 61—males; 50, 51, 53, 55, 58, 60, 62—females; 51—neotype; 54, 56, 57—holotypes; 58—paratype.

FIGURES 83–102. Apterodela spp., labrum: 83, 84, 88, 89—A. (s. str.) ovipennis (83, 88—Japan, Itoigawa city, Agero; 84, 89—Japan, Honsu, Mooka); 85–87, 90–92—A. (s. str.) lobipennis (85—China, Hubei, Chang Yang; 86—China, Hubei, Ichang; 87, 92—China, Hubei, Enshi; 90—China, Jiangxi, Kiu Kiang; 91—China, Hubei, Dabie Shan); 93, 98—A. (s. str.) kazantsevi (93—China, Jiangxi, Jinggangshan; 98—China, Jiangxi, Wufugang); 94, 99—A. (s. str.) latissima sp. nov. (China, Yunnan, Xishangbana); 95, 100—A. (s. str.) alopecomma sp. nov. (95—China, Sichuan, Qingcheng Hou Shan; 100—China, Sichuan, Guan Xian); 96, 101—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 97, 102—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P.); 83–87, 93–97, 99—males; 88–92, 98, 100–102—females; 90—neotype; 93–95—holotypes; 99, 100—paratypes.

FIGURES 123–142. Apterodela spp., pronotum: 123, 128,—A. (s. str.) ovipennis (123—Japan, Honsu, Mooka; 128—Japan, Itoigawa city, Agero); 124–127, 129–132—A. (s. str.) lobipennis (124—China, Hubei, Chang Yang; 125—China, Hubei, Ichang; 126—China, Kiang-Si [= Jiangxi]; 127, 132—China, Hubei, Enshi; 129—China, Jiangxi, Kiu Kiang; 130—China, Hubei, Dabie Shan; 131—China, Kiang-Si [= Jiangxi], Yao-chow); 133, 138—A. (s. str.) kazantsevi (133—China, Jiangxi, Jinggangshan; 138—China, Jiangxi, Wufugang); 134, 139—A. latissima sp. nov. (China, Yunnan, Xishangbana); 135, 140— A. (s. str.) alopecomma sp. nov. (135—China, Sichuan, Qingcheng Hou Shan; 140— China, Sichuan, Guan Xian); 136, 141—A. (Protopterodela) shirakii (China, Taiwan, Alishan); 137, 142—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P.); 123–127, 133–137, 139—males; 128–132, 138, 140–142—females; 133–135—holotypes; 139, 140—paratypes.

FIGURES 146–152. Apterodela spp., series umbilicata and metepisternum, right lateral view:146—A. (s. str.) ovipennis (Honsu, Mooka); 147—A. (s. str.) bivirgulata bivirgulata (China, Henan, Hui-Xian); 148—A. (s. str.) lobipennis (China, Hubei, Ichang); 149—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 150—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 151—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 152—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 146, 148–152—males; 147—female; 149, 150—holotypes. Without scales.

FIGURES 193–212. Apterodela (s. str.) spp., left elytron:193–198, 203–210—A. (s. str.) lobipennis (193—China, Hubei, Chang Yang; 194, 206—China, Hubei, Ichang; 195–197, 207, 208—China, Hubei, Enshi; 198—China, Kiang-Si [= Jiangxi], Yaochow; 203–205—China, Jiangxi, Kiu Kiang; 209—China, Hubei, Dabie Shan; 210—China, Zhejiang, Dapanshan); 199, 200— A. (s. str.) latissima sp. nov. (China, Yunnan, Xishangbana); 201, 211—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 202, 212—A. (s. str.) alopecomma sp. nov. (202—China, Sichuan, Qingcheng Hou Shan; 212—China, Sichuan, Guan Xian); 193–202—males; 203–212—females; 199, 201, 202—holotypes; 200, 212—paratypes; 203—neotype.

FIGURES 213–226. Apterodela spp., aedeagus, left lateral view: 213—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 214, 215—A. (s. str.) bivirgulata bivirgulata (214—China, Jiangxi, Lushan; 215—China, Zhejiang, Dao Shan Island); 216–219—A. (s. str.) bivitgulata occidentalis ssp. nov. (216 – China, Kansou, Hoei-Sien; 217—China, Shanxi, Zhong-Tiao-Shan; 218— China, Inner Mongolia, Hua-Shu-Ye; 219—China, Sichuan, Daheishan); 220, 221—A. (s. str.) lobipennis (220—China, Chang Yang; 221—China, Ichang); 222—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 223—A. (s. str.) latissima sp. nov. (China, Yunnan, Xishangbana); 224—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 225—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 226—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 216–219, 223—paratypes; 222, 224—holotype.

FIGURES 227–236. Apterodela spp., aedeagus, right lateral view: 227—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 228, 229—A. (s. str.) bivirgulata bivirgulata (228—China, Chin Kiang; 229—China, Zhejiang, Dao Shan Island); 230, 231—A. (s. str.) bivitgulata occidentalis ssp. nov.sp. nov. (230—China, Shanxi, Zhong-Tiao-Shan; 231—China, Inner Mongolia, Hua- Shu-Ye); 232—A. (s. str.) lobipennis (China, Ichang); 233—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 234—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 235—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 236—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 233—holotype; 230, 231, 234—paratypes.

FIGURES 237–243. Apterodela spp., internal sack, dorsal view: 237—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 238—A. (s. str.) bivirgulata bivirgulata (China, Chin Kiang); 239—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 240—A. (s. str.) lobipennis (China, Hubei, Enshi); 241—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 242—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 243—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 239, 241—paratypes. Without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 244–250. Apterodela spp., internal sac, left lateral view: 244—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 245— A. (s. str.) bivirgulata bivirgulata (China, Chin Kiang); 246—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 247—A. (s. str.) lobipennis (China, Hubei, Enshi); 248—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 249—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 250—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 246, 248—paratypes. Without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 251–257. Apterodela spp., internal sac, right lateral view: 251—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 252—A. (s. str.) bivirgulata bivirgulata (China, Chin Kiang); 253—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 254— A. (s. str.) lobipennis (China, Hubei, Enshi); 255—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 256—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 257—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 253, 255—paratypes. Without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 258–264. Apterodela spp., internal sack, ventral view: 258—A. (s. str.) ovipennis (Japan, Itoigawa, Agero); 259—A. (s. str.) bivirgulata bivirgulata (China, Chin Kiang); 260—A. (s. str.) kazantsevi (China, Jiangxi, Jinggangshan); 261—A. (s. str.) lobipennis (China, Hubei, Enshi); 262—A. (s. str.) alopecomma sp. nov. (China, Sichuan, Qingcheng Hou Shan); 263—A. (Protoapterodela) shirakii (China, Taiwan, Alishan); 264—A. (Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P., Greenbrier Cove); 260, 262—paratypes. Without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 153–172. Apterodela (s. str.) spp., left elytron: 153, 154, 163, 164—A. (s. str.) ovipennis (153, 163—Japan, Sado; 154, 164—Japan, Honsu, Mooka); 155–162, 165–172—A. (s. str.) bivirgulata bivirgulata (155—China, Ngan-Hoeï; 156, 157, 166, 167—China, Jiangsu, Chin-Kiang; 158—China, Kiangsu [= Jiangsu]; 159, 169, 170 China, Jiangsu, Nanjing; 160—China, Jiangxi, Lushan; 161—China, Hubei, Wuhan; 162, 172—China, Zhejiang, Dao Shan Island; 165—China, Shandong, Ho-Chan; 168—China, Henan, Hui-Xian; 171—China, Jiangsu, Hushukuan); 153–162—males; 163–172—females; 153—lectotype; 155—holotype; 163—paralectotype.

FIGURES 173–192. Apterodela spp., left elytron: 173–179, 183–189—A. (s. str.) bivirgulata occidentalis ssp. nov. (173— China, Kansu, eastern part of Kuku nor; 183, 184—China, Gansu, Hweisin; 174, 175, 185—China, Gansu, Hoei-Sienn; 176 – China, Henan, Donghe-cun; 177, 187—China, Shanxi, Zhong-Tiao-Shan; 178, 188—China, Inner Mongolia, Hua-Shu-Ye; 186—China, Gansu, Wangziguan; 179, 189—China, Sichuan, Daheishan); 180, 181, 190, 191—A. (Protoapterodela) shirakii (180—China,Taiwan,Tattka;190—China,Taiwan,Roeichi;181, 191—China,Taiwan,Alishan);182,192—A.(Protoapterodela) unipunctata (USA, TN, Great Smoky Mts N.P.); 173–182—males; 183–192—females; 183— holotypes; 173–175, 178, 179, 184–189—paratypes; 180—lectotype; 190—paralectotype.

FIGURES 273–278. Morphometric features of Apterodela spp.: 273–275—relation of the proportions in different species (273—relation of the proportions PW/PL and EW/EHW; 274, 275—relation of the proportions HW/PW and TL/BH; 274— males; 275—females; yellow rhombs—A. (Protoapterodela) unipunctata; red crosses—A. (Protoapterodela) shirakii; light green rectangles—A. (s. str.) ovipennis; lilac quadrates—A. (s. str.) bivirgulata; green triangles—A. (s. str.) lobipennis; blue circles—A. (s. str.) kazantsevi; orange rhombs—A. (s. str.) alopecomma sp. nov.; light blue asterisks—A. (s. str.) latissima sp. nov.; open symbols—females; solid symbols—males); 276–278—differences of the body proportions (276, 277—TL/BH; 278—HW/PW; m—males; f—females; small symbols in the rectangles—medians; rectangles—25–75 percentiles; whiskers— min–max; significantly different samples see in the Tables 1, 2).