Tomogenius papuaensis Gomy, 2007

Lackner, Tomas & Leschen, Richard A. B., 2017, A monograph of the Australopacific Saprininae (Coleoptera, Histeridae), ZooKeys 689, pp. 1-263 : 200-204

Tomogenius papuaensis Gomy, 2007 Figs 716, 717-722, 723-731, 755

Tomogenius papuaensis Gomy, 2007: 42, figs 2, 4, 6 and photographs.

Type locality.

Papua New Guinea: Tapini.

Type material examined.

Tomogenius papuaensis Gomy, 2007: paratype, ♂, side-mounted on triangular mounting card, with genitalia glued to another triangular mounting card situated below the specimen, with the following labels: "Nlle. GUINEE / Tapini VII. 1968 / (J. POULLARD réc.)” (printed); followed by: “♂” (printed); followed by: " Tomogenius / papuaensis Gomy / Y. Gomy - Det. 2009" (written-printed); followed by: "Y. Gomy des. / PARATYPE" (red label, printed); followed by: "Collection / Y. GOMY" (printed); followed by: “09-095” (yellow label, pencil-written, added by the senior author) (TLAN).

Biology.

Found in a cave in bat guano ( Gomy 2007).

Distribution.

Papua New Guinea: Tapini (Fig. 755).

Re-description.

Body length: PEL: 3.35 mm; APW: 1.20 mm; PPW: 2.25 mm; EL: 2.25 mm; EW: 2.75 mm. Body (Fig. 716) ovoid, moderately convex from above, underside slightly flattened, cuticle almost black without metallic luster; legs and body appendages castaneous brown. Antennal scape (Fig. 717) not particularly thickened, with few short setae; club rather large, oval, entirely covered in dense short sensilla, intermingled with sparse longer erect sensilla; sensory structures of antennal club not examined.

Mandibles with rounded outer margin, acutely pointed, labrum slightly convex, sparsely punctate; mentum sub-trapezoid, anterior angles slightly produced, anterior margin with a shallow median notch, surface around it with several long setae, disc without setae; other mouthparts not examined.

Clypeus (Fig. 717) large, rectangular, flattened, with sparse fine punctures separated by several times their diameter; frontal stria absent; supraorbital stria vaguely present; frontal disc (Fig. 717) with sparse fine round punctures larger and coarser than those of clypeus; eyes convex, well visible from above.

Pronotal sides (Fig. 716) on basal two-thirds feebly convergent anteriorly, on apical third strongly convergent, apical angles obtuse, marginal pronotal stria complete, thin, slightly carinate; disc medially covered with sparse round fine punctation, punctures separated by several times their diameter, laterally punctation becomes coarser and denser; pronotal hypomeron glabrous; scutellum very small.

Elytral epipleura with very fine scattered punctures; marginal epipleural stria double, both striae weakly impressed but complete; marginal elytral stria well impressed, continuous along elytral apex as apical elytral stria, stopping in middle of elytral apical margin; humeral elytral stria well impressed on basal fourth; inner subhumeral stria absent, short; elytral disc with four dorsal elytral striae 1-4, first the longest, reaching about two-thirds of elytral length apically, second and third striae ending short of elytral half, fourth dorsal elytral stria the shortest, reaching about elytral third apically; characteristic hooked appendix present between fourth dorsal elytral and sutural striae in other species of the genus absent; sutural elytral stria weakened, intermittent, erased on basal third and apical tenth. Entire elytral disc punctate, on basal half (roughly) punctures very fine and sparse, on apical half (roughly) punctures larger and denser, but still rather sparse, separated approximately two-three times their diameter.

Propygidium (Fig. 718) transverse, about four times as broad as long, completely exposed, with dense and coarse punctures separated by less than their diameter; pygidium (Fig. 718) with sparser punctation, becoming even sparser and finer towards apex.

Anterior margin of median portion of prosternum (Fig. 719) rounded; marginal prosternal stria present only laterally; prosternal process flattened, broad, sparsely punctate; carinal prosternal striae (Fig. 719) slightly bisinuate, terminating near large and deep united apical foveae; lateral prosternal striae carinate, slightly convergent anteriorly, attaining apices of carinal prosternal striae. Lateral costa of antennal groove not reaching prosternal process.

Anterior margin of mesoventrite (Fig. 720) almost straight; discal marginal mesoventral stria well impressed, slightly carinate; disc flattened, with coarse punctures separated by about their diameter; meso-metaventral suture vague, meso-metaventral sutural stria undulate, intermittent.

Intercoxal disc of metaventrite medially with slight longitudinal depression creating near basal margin two obtuse tubercles; disc of metaventrite rather coarsely punctate near anterior angles, medially punctures finer and sparser, becoming larger and coarser again along lateral margin. Lateral metaventral stria well impressed, carinate, shortened; lateral disc of metaventrite (Fig. 721) flattened, with round shallow large punctures fringed with microscopic setae; metepisternum + fused metepimeron (Fig. 721) evenly covered with much coarser and denser punctation, punctures without setae; lateral metepisternal stria present, deeply impressed and almost complete.

Intercoxal disc of first abdominal ventrite almost completely striate laterally; surface of disc with scattered oblong punctation, punctures becoming sparser and finer medially.

Protibia (Fig. 722) flattened and somewhat dilated, outer margin with six low teeth, topped by short triangular denticles followed by two-three minuscule denticles; setae of outer row short, sparse; setae of median row denser, slightly shorter than those of outer row; protarsal groove shallow; anterior protibial stria complete, costate; two thin, rather long tarsal denticles present apically; protibial spur short, straight, growing out from apical protibial margin; apical margin of protibia posteriorly with four tiny apical denticles; outer part of posterior surface of protibia finely imbricate, with a row of short setae; median part of posterior surface with additional row of minuscule setae; posterior protibial stria complete, with scattered minuscule setae turning into several tightly-spaced short and stout denticles near apical margin; inner margin with double row of short lamellate setae.

Mesotibia slender, outer margin with a single row of dense thin denticles growing in size apically; setae of outer row and those of median row not examined; posterior mesotibial stria not examined; anterior surface of mesotibia with dense row of well sclerotized short setae, with another similar row of much shorter situated below it; anterior mesotibial stria complete, terminating in numerous tiny inner anterior denticles; mesotibial spur broken off; apical margin with two tiny denticles; mesotarsus shorter than mesotibia; claws of apical tarsomere about half its length; metatibia basically similar to mesotibia, but denticles of outer margin much sparser than those of mesotibia; claws of apical tarsomere somewhat shorter, about one-third its length.

Male genitalia. Eighth sternite (Figs 723-724) fused medially; apically with a closely-set cluster of long, brush-like setae; eighth tergite and eighth sternite not fused laterally (Fig. 725). Ninth tergite (Figs 726-727) longitudinally divided medially; spiculum gastrale (Figs 728-729) gradually dilated in most of apical half, basal end dilated, spoon-like. Aedeagus (Figs 730-731) slender, subparallel, slightly bisinuate before apex; basal piece of aedeagus short, ratio of its length: length of parameres 1: 4; parameres fused along their basal two-thirds (roughly); aedeagus slightly curved from lateral view (Fig. 731).