Molycria mammosa, : Simon, 1897

Platnick, N. I. & Baehr, B., 2006, A Revision Of The Australasian Ground Spiders Of The Family Prodidomidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2006 (298), pp. 1-287 : 30-35

Molycria mammosa (O. P.-Cambridge) Figures 61–65; Map 4

Mutusca mammosa O. P.- Cambridge, 1874: 173, pl. 17, fig. 2 (male holotype from ‘‘Shelley’s Flats,’’ New South Wales, Australia, in HDO, examined).

Molycria mammosa: Simon, 1897: 158 . — Deeleman-Reinhold, 2001: 555, figs. 953–957.

DIAGNOSIS: Males resemble those of M. smithae but have a much flatter conductor tip (fig. 62); females also resemble those of M. smithae but lack the pronounced, posteriorly directed internal lobes on the posterior margin of the epigynal atrium that are characteristic of that species, having ducts that instead grade gently into the posterior atrial margins (fig. 65).

MALE: Total length 3.24. Carapace 1.50 long, 1.44 wide, 0.42 high, length/width 1.04; sternum 0.98 long, 0.82 wide, length/width 1.19; abdomen 1.74 long, 0.96 wide; coxa I 0.56 long; relative length of coxae I–IV 1.00:0.71:0.68:0.96. Carapace, sternum, chelicerae, legs grayish orange; endites, labium orange, distally pale; abdominal dorsum gray, with weak orange scutum, spear tipshaped pale spot in front of spinnerets, venter pale. Carapace weakly covered with gray, plumose setae. AME elevated, eye group width 0.77 of caput width; AME 0.16; ALE 0.11; PME 0.14; PLE 0.11; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.04; PME– PLE 0.06; ALE–PLE 0.02; eye group AME– PME 0.34; AME–AME 0.34; PME–PME 0.32. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.53 of abdominal length, contiguous. Palp (figs. 61– 63): conductor originating distally, sclerotized, grooved longitudinally, with ventrally folded tip; median apophysis wide, ventrally excavated, with two distal tips, median one bifurcate; sperm duct u-shaped; embolus long, straight; retrolateral tibial apophysis long, triangular, dorsally directed.

FEMALE: Total length 3.50. Carapace 1.50 long, 1.44 wide, 0.42 high, length/width 1.04; sternum 0.96 long, 0.88 wide, length/width 1.09; abdomen 2.00 long, 1.20 wide; relative length of coxae I–IV 1.00:0.78:

0.75:1.00. Coloration as in male but without dorsal scutum. Eye group width 0.73 of caput width; AME 0.15; PME 0.14; PME–PME 0.02; eye group AME–PME 0.30; AME–AME 0.32; PME–PME 0.30. Clypeus 0.08 high. ALS 0.48 of abdominal length. Epigynum (figs. 64, 65): atrium broadly rectangular, with arched posterior margin; epigynal ducts extremely short, in x-shaped position with spermathecae, spermathecae about their diameter apart, oval, twisted, with one dorsal diverticulum.

MATERIAL EXAMINED: Australian Capital Territory: Lees and Blundells Creeks, Brindabella Ranges , 35 ° 229S, 148 ° 509E, Oct. 10–Feb. 29, 1980 –1981, pitfalls (C. Dickman, ANIC), 7 Oi, 5♀ ; Piccadilly Circus , 35 ° 229S, 148 ° 489E, Jan. 1984, elev. 1240 m (J. Lawrence, T. Weir, M. Johnson, ANIC), 2 Oi ; Tidbinbilla Nature Reserve , 35 ° 289S, 148 ° 529E, Mar. 9, 1978 (P. Ormay, AMS KS13882 ), 1♀ ; Wombat Creek, 6 km NE Piccadilly Circus, 35 ° 199S, 148 ° 519E, Feb. 1984, elev. 750 m (J. Lawrence, T. Weir, M. Johnson, ANIC), 1♀. New South Wales: Arara East State Forest , 10 km N Coff’s Harbour, ca. 30 ° 189S, 153 ° 079E, May 18, 1979, under log (D. Hirst, SAM NN22281 ), 1 Oi ; 2 km S Ballengarra Creek crossing on Grey’s Road junction, Ballengarra State Forest , 31 ° 149S, 152 ° 459E, Feb. 5–Apr. 9, 1993, pitfall, elev. 120 m (M. Gray, G. Cassis, AMS KS43229 ), 1♀ ; Beecroft , 33 ° 459S, 151 ° 049E, Nov. 20, 1992 (J. Noble, AMS KS55259 ), 1 Oi, Nov. 22, 1993 (L. Maher, AMS KS57341 ), 1♀, June 16, 1994 (J. Noble, AMS KS53685 ), 1 Oi, June 24, 2001 (J. Noble, AMS KS76816 ), 1♀, July 4–25, 2001 –2002 (J. Noble, AMS KS76827 , 79752 ), 2Oi ; Booti Booti National Park, 32 ° 149S, 152 ° 339E, Dec. 14, 1996, pitfall (L. Wilkie, AMS KS68359 ), 1 Oi , Nov. 25, 1997, pitfall (L. Wilkie, AMS KS60355 ), 1 Oi ; Bulls Ground State Forest, near Wauchope , 31 ° 339S, 152 ° 389E, Feb. 21–25, 1999, pitfall (A. York, AMS KS57642 ), 1♀ ; Bungawalbin State Forest , 29 ° 049S, 153 ° 079E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73953 ), 1 Oi ; Bungawalbin State Forest , 29 ° 059S, 153 ° 109E, Feb. 1997, eucalypt/paperbark forest pitfall (A. York, AMS KS73954 ), 1 Oi ; ridge between Camp Creek and Stydgy Creek, Leasehold land, 28 ° 469S, 152 ° 189E, Feb. 4–Apr. 9, 1993, pitfall, elev. 640 m (M. Gray, G. Cassis, AMS KS37081 ), 2 Oi ; Cooperabung Range Road, c. 1.1 km S Narang Road, Ballengarra State Forest , 31 ° 129S, 152 ° 429E, Feb. 4–Apr. 9, 1993, pitfall, elev. 200 m (M. Gray, G. Cassis, AMS KS43228 ), 2♀ ; 0.3 km S along Coxs Fence Trail from Boundary Road, Nerong State Forest , 31 ° 389S, 152 ° 099E, Feb. 5–Apr. 9, 1993, pitfall (M. Gray, G. Cassis, AMS KS43220 ), 1 Oi, 1♀ ; East Kunderang Trail , 2.1 km E West Kunderang Trail, 30 ° 499S, 152 ° 039E, Feb. 4–Apr. 9, 1993, pitfall, elev. 845 m (M. Gray, G. Cassis, AMS KS43212 ), 1♀ ; Enfield , 33 ° 549S, 151 ° 069E ( AMS KS17277 ), 1 Oi ; 1.2 km up track N Grey Gums Forest Road, 2.1 km from Doyles River Road, Bulga State Forest , 31 ° 329S, 152 ° 149E, Feb. 4–Apr. 9, 1993, pitfall, elev. 620 m (M. Gray, G. Cassis, AMS KS43235 ), 1♀ ; Homewoods Road, 2.8 km W Knodingbul Road, Bulga State Forest , 31 ° 379S, 152 ° 079E, Feb. 4–Apr. 9, 1993, pitfall, elev. 690 m (M. Gray, G. Cassis, AMS KS43231 ), 3♀ ; 600 m N Homewoods Road, 2.8 km W Knodingbul Road, Bulga State Forest , 31 ° 379S, 152 ° 079E, Feb. 4–Apr. 9, 1993, pitfall, elev. 650 m (M. Gray, G. Cassis, AMS KS43232 ), 1♀ ; Hornsby, Waitara Creek , 33 ° 429S, 151 ° 069E, Nov. 2001 (G. Milledge, H. Smith, AMS KS74990 ), 1 Oi ; Horseshoe Road, ca. 0.5 km N Scotchman Peak, Diehappy State Forest , 30 ° 299S, 152 ° 409E, Nov. 11–24, 1999, pitfall (M. Gray, G. Milledge, H. Smith, AMS KS61692 ), 1♀ ; Jamieson Park, Narrabeen, 33 ° 439S, 151 ° 189E, Nov. 1995, pitfall, elev. 40 m, ridge top Angophora costata woodland (M. Gray, H. Smith , AMS KS49021 ), 1 Oi , Nov. 6–20, 1995, same except elev. 20 m (M. Gray, H. Smith, AMS KS49013 ), 1♀, Mar. 5–19, 1996, same (M. Gray, H. Smith, AMS KS49053 ), 1♀ ; Jenolan , 33 ° 499S, 150 ° 029E, Dec. 1979, in house ( AMS KS9967 ), 1 Oi ; 2.3 km N Karuah River on Karuah River Road, Chichester State Forest , 32 ° 059S, 151 ° 439E, Feb. 4–Apr. 9, 1993, pitfall, elev. 520 m (M. Gray, G. Cassis, AMS KS38896 ), 1♀ ; Linton Nature Reserve , 350 m S road, on slope above small gulley, ca. 700 m W of reserve entrance, 30 ° 289S, 150 ° 539E, Nov. 18–Dec. 9, 2001, pitfall (H. Doherty, M. Elliott, AMS KS82216 ), 1 Oi, 1♀ ; junction, Moonpar and Mills Roads, Moonpar State Forest , 30 ° 159S, 152 ° 389E, Nov. 9–23, 1999, pitfall (M. Gray, G. Milledge, H. Smith, AMS KS61720 ), 1♀ ; 4.5 km NE from creek crossing, Morgans Creek, on Wheatly Creek Access Road, Leasehold land, 28 ° 469S, 152 ° 189E, Feb. 4–Apr. 9, 1993, pitfall, elev. 590 m (M. Gray, G. Cassis, AMS KS37030 ), 2 Oi, 1♀ ; 5.9 km NE from creek crossing on Morgans Creek, Leasehold land, 28 ° 469S, 152 ° 189E, Feb. 4–Apr. 9, 1993, pitfall, elev. 620 m (M. Gray, G. Cassis, AMS KS37083 ), 2 Oi, 1♀ ; Mount Allyn Road, 2.3 km N Shellbrook Forest Road, Chichester State Forest , 32 ° 089S, 151 ° 279E, Feb. 4–Apr. 9, 1993, pitfall, elev. 770 m (M. Gray, G. Cassis, AMS KS43213 ), 1♀ ; Mount Boss State Forest , 31 ° 129S, 152 ° 249E, Oct. 1980 ( AMS KS42863 , 43562 ), 1 Oi, 1♀ ; Mount Brown Road, 0.1 km N from Y intersection at Mount Brown, Richmond Range State Forest , 28 ° 379S, 152 ° 439E, Feb. 4–Apr. 9, 1993, pitfall, elev. 480 m (M. Gray, G. Cassis, AMS KS36024 ), 2 Oi, 1♀ ; Mummel Forest Road, 7.6 km N junction with Enfield Forest Road, Enfield State Forest , 31 ° 179S, 151 ° 519E, Feb. 4–Apr. 9, 1993, elev. 1340 m (M. Gray, G. Cassis, AMS KS41069 ), 1♀ ; Myall Lakes National Park , 32 ° 299S, 152 ° 249E, Oct. 10, 1997, pitfall (L. Wilkie, AMS KS60398 , 60401 ), 2 Oi ; Myrtle State Forest , 29 ° 119S, 152 ° 599E, Feb. 1997, eucalypt/paperbark forest pitfall (A. York, AMS KS73947 , 73955 ), 1 Oi, 2♀ ; Narraweena , 33 ° 459S, 151 ° 169E, July 25, 1997 (M. Robinson, AMS KS55253 ), 1 Oi ; Northern Fire Trail, Monga State Forest , 35 ° 339S, 149 ° 539E, Mar. 16, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63894 ), 1 Oi ; Oaky Creek Road, Richmond Range State Forest , 28 ° 389S, 152 ° 459E, Feb. 4– Apr. 9, 1993, pitfall, elev. 230 m (M. Gray, G. Cassis, AMS KS36053 ), 2 Oi, 2♀ ; Reids Creek Road, Gladstone State Forest , 30 ° 319S, 152 ° 489E, Nov. 12–25, 1999, pitfall (M. Gray, G. Milledge, H. Smith, AMS KS61099 ), 1 Oi ; Rivertree Fire Trail, on ridge ca. 2 km NNE from turnoff, Gilgurry State Forest , 28 ° 459S, 152 ° 159E, Feb. 4–Apr. 9, 1993, pitfall, elev. 770 m (M. Gray, G. Cassis, AMS KS37082 ), 1♀ ; 1.9 km W along Sheas Nob Road from Boundary Creek Road, Boundary Creek State Forest , 29 ° 599S, 152 ° 349E, Feb. 4–Apr. 9, 1993, pitfall, elev. 550 m (M. Gray, G. Cassis, AMS KS43216 ), 1♀ ; ‘‘ Shelley’s Flats’ ’ [5Shelleys Flat], 34 ° 449S, 149 ° 569E, 1874 (H. Bradley, HDO), 1 Oi (holotype) ; Stewarts Brook State Forest , 31 ° 549S, 151 ° 239E, Feb. 4–Apr. 9, 1993, pitfall (M. Gray, G. Cassis, AMS KS43154 ), 1 Oi ; Turtle Creek Fire Trail, Monga State Forest , 35 ° 389S, 149 ° 569E, Mar. 15, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63906 ), 1♀ ; junction, Wattle Creek Road and Wattle Creek, Richmond Range State Forest , 28 ° 389S, 152 ° 469E, Feb. 4–Apr. 9, 1993, pitfall, elev. 130 m (M. Gray, G. Cassis, AMS KS36040 ), 1 Oi, 2♀ ; 2.8 km from Wheatly Creek Access Road on Camp Creek, Leasehold land, 28 ° 479S, 152 ° 189E, Feb. 4–Apr. 9, 1993, pitfall, elev. 435 m (M. Gray, G. Cassis, AMS KS37085 ), 2♀ ; 0.5 km from Wheatly Creek Road on Camp Creek Road, Leasehold land, 28 ° 479S, 152 ° 199E, Feb. 4–Apr. 9, 1993, pitfall, elev. 550 m (M. Gray, G. Cassis, AMS KS37084 ), 2 Oi ; headwaters, Wheatleys Creek, Leasehold land, 28 ° 459S, 152 ° 199E, Feb. 4–Apr. 9, 1993, pitfall, elev. 550 m (M. Gray, G. Cassis, AMS KS37087 ), 5 Oi, 8♀ ; Wild Cattle Creek, 400 m NE Spur Track, Bulga State Forest , 31 ° 369S, 152 ° 079E, Feb. 4–Apr. 9, 1993, pitfall, elev. 400 m (M. Gray, G. Cassis, AMSKS43233, 43234), 2♀ .

DISTRIBUTION: Widely distributed in New South Wales and the Australian Capital Territory (map 4).

Molycria smithae , new species

Figures 66–70; Map 5

TYPES: Male holotype and female allotype from Brou Lake Road, Narooma , 36 ° 079S, 150 ° 039E, New South Wales (Mar. 9, 1999; L. Wilkie, R. Harris, H. Smith), deposited in AMS (male KS63896, female KS90086) .

ETYMOLOGY: The specific name is a patronym in honor of Helen Smith, of the Australian Museum, the collector of this species and many other interesting prodidomids.

DIAGNOSIS: Males resemble those of M. mammosa but have two distally directed lobes on the tip of the palpal conductor, separated by a sharply pointed, retrolaterally directed ridge (fig. 67); females have distinctive, heavily sclerotized, posteriorly directed internal protuberances at the sides of the epigynal atrium, overlying (in dorsal view) the epigynal ducts (fig. 70).

MALE: Total length 3.28. Carapace 1.48 long, 1.30 wide, 0.50 high, length/width 1.14; sternum 0.84 long, 0.82 wide, length/width 1.02; abdomen 1.80 long, 0.88 wide; coxa I 0.54 long; relative length of coxae I–IV 1.00: 0.78:0.70:0.92. Carapace, sternum, chelicerae, legs grayish orange; endites, labium orange, distally pale; abdominal dorsum gray, with weak orange scutum, half moon-shaped pale spot in front of spinnerets, venter pale, epigastric area orange. Carapace weakly covered with gray, plumose setae. Eye group width 0.73 of caput width; AME 0.14; ALE 0.10; PME 0.15; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.03; PME– PLE 0.02; ALE–PLE 0.04; eye group AME– PME 0.32; AME–AME 0.31; PME–PME 0.32. Clypeus 0.12 high. Abdomen covered with gray, plumose setae; ALS 0.49 of abdominal length, slightly separated. Palp (figs. 66– 68): conductor originating distally, with ventrally folded tip and two distally directed lobes, separated by sharply pointed, retrolaterally directed ridge; median apophysis wide, ventrally excavated, with two distal tips, median one bifurcate; sperm duct weakly u-shaped; embolus long, with ventrally folded distal end

68. Same, retrolateral view. 69. Epigynum, ventral view. 70. Same, dorsal view.

and retrolaterally bent tip; retrolateral tibial apophysis long, triangular, dorsally directed.

FEMALE: Total length 4.06. Carapace 1.58 long, 1.50 wide, 0.56 high, length/width 1.05; sternum 1.00 long, 0.92 wide, length/width 1.07; abdomen 2.48 long, 1.40 wide; coxa I 0.62 long; relative length of coxae I–IV 1.00:0.80:0.71:0.97. Coloration as in male but without scutum. Eye group width 0.77 of caput width; AME 0.15; ALE 0.14; PME 0.15; PLE 0.12; AME–AME 0.04; PME–PME 0.04; AME–AME 0.34; PME–PME 0.34. Clypeus 0.10 high. ALS 0.48 of abdominal length. Epigynum (figs. 69, 70): atrium broadly rectangular, with arched posterior margin; epigynal ducts extremely short, in x-shaped position with spermathecae, spermathecae less than their diameter apart, oval, twisted, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: New South Wales: Benandarah State Forest, near Benandarah , 35 ° 389S, 150 ° 139E, Dec. 11, 1977 – Feb. 18, 1978, pitfall, moist sclerophyll forest (M. Gray, AMS KS1250 ), 1♀ ; Benandarah State Forest , 8 km N Batemans Bay, 35 ° 409S, 150 ° 149E, Nov. 30, 1978 – Jan. 4, 1979, litter pitfall (C. Horseman, AMS KS2278 ), 1 Oi ; Blue Mountains, road to Ingar Picnic Area , 33 ° 469S, 150 ° 259E, Oct. 3, 1996, pitfall ( AMS KS55254 ), 1♀ ; Blue Mountains, road to Ingar Picnic Area , 33 ° 469S, 150 ° 269E, Oct. 2, 1996, pitfall ( AMS KS55256 ), 1 Oi ; 0.45 km E along Boundary Road from Masonite Track, Nerong State Forest , 32 ° 379S, 152 ° 079E, Feb. 4–Apr. 9, 1963, pitfall, elev. 130 m (M. Gray, G. Cassis, AMS KS43219 ), 1♀ ; Bullaburra, Red Gum Park , 33 ° 449S, 150 ° 259E, Oct. 4, 1996, pitfall ( AMS KS55257 ), 1♀ ; junction, Carls Mountain and North Head Roads, Murramarang National Park , 35 ° 419S, 150 ° 169E, Mar. 17, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63908 ), 1♀ ; Coomerang Road, Dampier State Forest , 36 ° 049S, 149 ° 559E, Mar. 11, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63902 ), 3♀ ; Coomerang Road, Dampier State Forest , 36 ° 079S, 149 ° 599E, Mar. 10, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63911 ), 1♀ ; Fire Trail , 2.5 km WSW Durras, 35 ° 409S, 150 ° 169E, Mar. 17, 1999 (J. Tarnawski, S. Lassau, AMS KS63899 ), 2♀ ; Forest Drive, Kioloa State Forest , 15 km N Batemans Bay, 35 ° 369S, 150 ° 169E, Sept. 7–Oct. 4, 1979, litter pitfall (C. Horseman, AMS KS3905 ), 1♀ ; Forest Drive, Kioloa State Forest, 18 km N Batemans Bay, 35 ° 379S, 150 ° 169E, Sept. 7–Jan. 4, 1978 –1979, litter pitfall (C. Horseman, AMS KS2128 , 2299 , 2812 ), 3 Oi , Jan. 4–30, 1979, litter pitfall (C. Horseman, AMS KS2475 ), 1♀ ; Gilmore Lookout, Wallaroo State Forest , 32 ° 379S, 151 ° 479E, Feb. 4–Apr. 9, 1963, pitfall, elev. 210 m (M. Gray, G. Cassis, AMS KS43217 , 43218 ), 3♀ ; Gosford-Ourimbah , 33 ° 249S, 151 ° 219E, May 1993, rainforest, under logs ( AMS KS35168 ), 1♀ ; Highway 54, 32 km NW Batemans Bay , 35 ° 339S, 150 ° 009E, Mar. 16, 1999 (J. Tarnawski, S. Lassau, AMS KS63912 ), 1♀ ; Jabarrah Point walking trail, Bodalla State Forest , 36 ° 059S, 150 ° 089E, Mar. 9, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63910 ), 1♀ ; 7.5 km S Nelligen, Mogo State Forest , 35 ° 439S, 150 ° 079E, Mar. 8, 1999 (J. Tarnawski, S. Lassau, AMS KS63901 ), 1♀ ; No Name Fire Trail, Buckenbowra State Forest , 35 ° 389S, 150 ° 009E, Mar. 15, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63909 ), 1♀ ; North Head Road, Murramarang National Park , 35 ° 429S, 150 ° 179E, Mar. 17, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63907 ), 1♀ ; Orange Ridge Road, Bodalla State Forest , 36 ° 179S, 149 ° 549E, Mar. 12, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63897 ), 1 Oi, 4♀ ; 1.5 km W Potato Point, Bodalla State Forest , 36 ° 069S, 150 ° 079E, Mar. 9, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63900 ), 1♀ ; Princess Highway, 1.5 km S Bodalla, Bodalla State Forest , 36 ° 079S, 150 ° 039E, Mar. 9, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63898 ), 2♀ ; Richmond Beach Road, Murramarang National Park , 35 ° 419S, 150 ° 179E, Mar. 17, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63903 ), 1♀ ; Ross Rixon Road, Dampier State Forest , 36 ° 039S, 149 ° 589E, Mar. 11, 1999 (L. Wilkie, R. Harris, H. Smith, AMS KS63904 ), 5♀ ; junction, Windywoppa and Lookout Roads, Boyne State Forest , 35 ° 369S, 150 ° 129E, Mar. 17, 1999 (J. Tarnawski, S. Lassau, AMS KS63895 ), 1 Oi ; Wiola Creek Fire Trail, Badja State Forest , 36 ° 079S, 149 ° 379E, Mar. 13, 1999 (J. Tarnawski, S. Lassau, AMS KS63905 ), 2♀ ; Woronora Dam Catchment, Princess Highway , ca. 0.1 km NW Southern Freeway Overpass, 34 ° 129S, 150 ° 589E, Dec. 8– 22, 1999, pitfall (M. Gray, G. Milledge, H. Smith, AMS KS63241 ), 1♀ ; Wyrrabalong National Park, 33 ° 179S, 151 ° 339E, June 2, 1997 (L. Wilkie, AMS KS63685 ), 1 Oi , Oct. 11, 1997, pitfall (L. Wilkie, AMS KS60363 ), 1♀, Nov. 27, 1997, pitfall (L. Wilkie, AMS KS60357 ), 1 Oi .

DISTRIBUTION: Known only from New South Wales (map 5).