Molycria milledgei, Platnick & Baehr, 2006

Platnick, N. I. & Baehr, B., 2006, A Revision Of The Australasian Ground Spiders Of The Family Prodidomidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2006 (298), pp. 1-287 : 35-41

Molycria milledgei , new species Figures 71–75; Map 5

TYPES: Male holotype and female allotype taken in pitfall trap in SW corner of Linton Nature Reserve , 60 m E from road, 30 ° 289S, 150 ° 529E, New South Wales (Nov. 18–Dec. 9, 2001; H. Doherty, M. Elliott) ,

deposited in AMS (male KS82209, female KS90087).

ETYMOLOGY: The specific name is a patronym in honor of Graham Milledge, the collection manager for arachnids at the Australian Museum, who provided excellent technical assistance.

DIAGNOSIS: Males can be recognized by the wide embolus and the squared protuberance on the base of the conductor (fig. 72), females by the long epigynal margins (fig. 74).

MALE: Total length 4.06. Carapace 1.76 long, 1.58 wide, 0.48 high, length/width 1.11; sternum 1.02 long, 0.90 wide, length/width 1.13; abdomen 2.30 long, 1.16 wide; coxa I 0.70 long; relative length of coxae I–IV 1.00:0.83:0.60:0.94. Carapace, sternum, chelicerae, legs grayish orange; endites, labium grayish, distally pale; abdominal dorsum gray, with weak orange scutum, half ringshaped pale spot in front of spinnerets, venter pale, epigastric area orange. Carapace weakly covered with gray, plumose setae. AME elevated, eye group width 0.79 of caput width; AME 0.16; ALE 0.14; PME 0.15; PLE 0.14; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.04; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.36; AME–AME 0.34; PME–PME 0.34. Clypeus 0.12 high. Abdomen covered with gray, plumose setae; ALS 0.54 of abdominal length, slightly separated. Palp (figs. 71–73): conductor originating distally, with ventrally folded distal end, retrolaterally directed, beak-shaped tip, squared protuberance on base; median apophysis about twice as long as wide, ventrally excavated, with two distal tips; sperm duct weakly u-shaped; embolus long, straight, with retrolaterally bent tip; retrolateral tibial apophysis long, triangular, dorsally directed.

FEMALE: Total length 4.14. Carapace 1.58 long, 1.54 wide, 0.42 high, length/width 1.02; sternum 0.98 long, 1.06 wide, length/ width 2.56; abdomen 1.32 long, 0.100 wide; coxa I 0.62 long; relative length of coxae I–IV 1.00:0.83:0.74:0.96. Coloration as in male but without scutum. Eye group width 0.74 of caput width; AME 0.16; ALE 0.12; PME 0.15; PLE 0.12; eye group AME–PME 0.30. Clypeus 0.10 high. ALS 0.48 of abdominal length. Epigynum (figs. 74, 75): atrium with rounded anterior margin, arched posterior margin; epigynal ducts short, in x-shaped position with spermathecae, spermathecae less than their diameter apart, oval, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: New South Wales: Attunga State Forest , SE side, E road up slope, 30 ° 579S, 150 ° 569E, Nov. 16–Dec. 7, 2001, pitfall (G. Carter, AMS KS82217 ), 1 Oi ; Linton Nature Reserve, 300 m from reserve entrance from Warranbah , 800 m S road, 30 ° 289S, 150 ° 549E, Nov. 18–Dec. 9, 2001, pitfall (H. Doherty, M. Elliott, AMS KS82215 ), 1♀ ; Mount Kaputar National Park, ca. 50 m NE picnic area at end of Bullawa Creek Road, 30 ° 149S, 150 ° 079E, Nov. 20–Dec. 11, 2001, pitfall (L. Wilkie, H. Smith, AMS KS82210– 82212 ), 2 Oi, 1♀ ; Mount Kaputar National Park, N side Mount Coryah, 200 m S road to Mount Kaputar , 30 ° 179S, 150 ° 079E, Nov. 20–Dec. 11, 2001, pitfall (H. Doherty, M. Elliot, AMS KS82213 , 82214 ), 1 Oi, 1♀ .

DISTRIBUTION: Known only from New South Wales (map 5).

Molycria grayi , new species

Figures 76–80; Map 5

TYPES: Male holotype and female allotype taken in pitfall traps on trail to Intermediate Hill, North Hummock, Lord Howe Island, 31 ° 339S, 159 ° 059E, New South Wales (Nov. 26–Dec. 3, 2000), deposited in AMS (male KS90088, female KS90089).

ETYMOLOGY: The specific name is a patronym in honor of Dr. Michael Gray of the Australian Museum, in recognition of his important works on Australian spiders.

DIAGNOSIS: Males resemble those of M. smithae , but have a single distally produced lobe on the prolateral side of the palpal conductor (fig. 77); females have the epigynal ducts restricted to the median portion of the epigynum (fig. 80).

MALE: Total length 3.46. Carapace 1.62 long, 1.46 wide, 0.44 high, length/width 1.11; sternum 0.88 long, 0.86 wide, length/width 1.02; abdomen 1.84 long, 1.00 wide; coxa I 0.56 long; relative length of coxae I–IV 1.00:0.79:0.71:0.92. Carapace, sternum, chelicerae, legs grayish orange; endites, labium orange brown, distally pale; abdominal dorsum gray, with weak orange scutum, spear tip-shaped pale spot in front of spinnerets, venter pale. Carapace weakly covered with gray, plumose setae. AME elevated, eye group width 0.79 of caput width; AME 0.16; ALE 0.10; PME 0.14; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME–PME 0.06; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.34; AME–AME 0.34; PME–PME 0.34. Clypeus 0.14 high. Abdomen covered with gray, plumose setae; ALS 0.55 of abdominal length, contiguous. Palp (figs. 76–78): conductor originating distally, with ventrally folded distal end bearing single distally produced lobe; median apophysis about twice as long as wide, ventrally excavated, with two 78. Same, retrolateral view. 79. Epigynum, ventral view. 80. Same, dorsal view.

distal tips; sperm duct weakly u-shaped; embolus long, with bifid tip, prolateral part bent retrolaterally; retrolateral tibial apophysis short, dorsally directed.

FEMALE: Total length 3.58. Carapace 1.54 long, 1.50 wide, 0.48 high, length/width 1.03; sternum 0.98 long, 0.88 wide, length/ width 1.11; abdomen 2.04 long, 1.06 wide; coxa I 0.58 long; relative length of coxae I–IV 1.00:0.76:0.72:0.96. Coloration as in male but without scutum. AME 0.15; PME 0.14; PLE 0.12; PME–PME 0.04; AME–AME 0.32; PME–PME 0.32. Clypeus 0.10 high. ALS 0.61 of abdominal length. Epigynum (figs. 79, 80): atrium broadly rectangular, with arched posterior margin; epigynal ducts short, in x-shaped position with spermathecae, spermathecae less than their diameter apart, oval, twisted, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: New South Wales: 25 m above coastal trail to Boat Harbour, 750 m from start, Lord Howe Island , 31 ° 339S, 159 ° 059E, Dec. 3–13, 2000, pitfall ( AMS KS76095 ), 1♀ ; 300 m NW Gilgurry Mountain, Boorook State Forest , 28 ° 479S, 152 ° 109E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 980 m (M. Gray, G. Cassis, AMS KS37086 ), 1♀ ; Goanna Creek Road, Richmond Range State Forest , 28 ° 369S, 152 ° 419E, Feb. 4–Apr. 9, 1993, pitfall, elev. 545 m (M. Gray, G. Cassis, AMS KS36018 ), 3♀ ; Main Creek, track off Mount Tindal Road, Ramornie State Forest , 29 ° 439S, 152 ° 389E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 110 m (M. Gray, G. Cassis, AMS KS43222 , 43223 ), 2 Oi, 1♀ ; 5.9 km NE from creek crossing on Morgans Creek, Leasehold land, 28 ° 469S, 152 ° 189E, Feb. 4–Apr. 9, 1993, pitfall, elev. 620 m (M. Gray, G. Cassis, AMS KS37088 ), 1♀ ; Mount Tindal Road, Ramornie State Forest , 29 ° 419S, 152 ° 359E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 490 m (M. Gray, G. Cassis, AMS KS43225 ), 2♀ ; Mount Tindal Road, Ramornie State Forest , 29 ° 429S, 152 ° 359E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 380 m (M. Gray, G. Cassis, AMS KS43227 ), 1♀ ; track off Mount Tindal Road, Ramornie State Forest , 29 ° 429S, 152 ° 379E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 220 m (M. Gray, G. Cassis, AMS KS43226 ), 2♀ ; North Hummock, trail to Intermediate Hill, Lord Howe Island , 31 ° 339S, 159 ° 059E, Nov. 26–Dec. 3, 2000, pitfall ( AMS KS75973 ), 1 Oi , Feb. 18–27, 2001, pitfall ( AMS KS79130 ), 1 Oi ; Peach Tree Ridge, just below summit of Intermediate Hill, Lord Howe Island , 31 ° 339S, 159 ° 059E, Nov. 26–Dec. 3, 2000, pitfall ( AMS KS76030 ), 1 Oi , Dec. 3–13, 2000, pitfall ( AMS KS76031 ), 1 Oi , Feb. 18–27, 2001, pitfall ( AMS KS79131 ), 1♀ ; Stephens Reserve, New Settlement, Lord Howe Island , 31 ° 329S, 159 ° 049E, Dec. 4–14, 2000, pitfall ( AMS KS76252 ), 1 Oi ; Stevens Reserve, Lord Howe Island , 31 ° 329S, 159 ° 049E, Feb. 25, 2001, fogging greybark and maulwood trunks (G. Milledge, AMS KS70599 ), 1 Oi ; SE face, Transit Hill, near summit, Lord Howe Island, 31 ° 329S, 159 ° 059E, Nov. 24–Dec. 1, 2000, pitfall ( AMS KS75958 , 75961 , 75964 , 75968 ), 4 Oi , Dec. 1–11, 2000, pitfall ( AMS KS75971 ), 1♀ ; W slope, Transit Hill, Lord Howe Island , 31 ° 329S, 159 ° 029E, Nov. 24–Dec. 1, 2000, pitfall ( AMS KS75940 , 75943 , 75952 ), 1 Oi, 4♀ ; T-Ridge Road, Ramornie State Forest , 29 ° 439S, 152 ° 339E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 300 m (M. Gray, G. Cassis, AMS KS43224 ), 2♀ ; Tributary Grasstree Creek, Nogrigar Road, Ewingar State Forest , 29 ° 089S, 152 ° 259E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 720 m (M. Gray, G. Cassis, AMS KS43214 ), 1♀ ; headwaters, Valorem Creek, track off Mount Tindal Road, Ramornie State Forest , 29 ° 429S, 152 ° 359E, Feb. 4–Apr. 9, 1963, pitfalls, elev. 380 m (M. Gray, G. Cassis, AMS KS43221 ), 2 Oi, 1♀. Queensland: Stanthorpe, 28 ° 399S, 151 ° 569E, Aug. 15, 1987, woodland (W. McKenzie, QMB S26410 ), 1Oi .

DISTRIBUTION: Southeastern Queensland and adjacent New South Wales, and also found on Lord Howe Island (map 5).

Molycria bundjalung , new species

Figures 81–85; Map 6

TYPES: Male holotype and female allotype taken in pitfall traps on hilltop on ridge between Big Marsh and Esk River, Bundjalung National Park, 29 ° 179S, 153 ° 169E, New South Wales (Feb. 4–Apr. 9, 1963; M. Gray, G. Cassis), deposited in AMS (male KS43236, female KS90487) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males can easily be distinguished from those of the other mammosa group species by the distally bent tibial apophysis (fig. 83), females by the very wide epigynal atrium, with ducts extending for its full width (figs. 84, 85).

MALE: Total length 3.42. Carapace 1.58 long, 1.42 wide, 0.54 high, length/width 1.11; sternum 0.90 long, 0.82 wide, length/width 1.09; abdomen 1.84 long, 0.94 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.84:0.76:0.96. Carapace, sternum, chelicerae, legs grayish orange; endites, labium orange brown, distally pale; abdominal dorsum gray, with weak orange scutum, spear tip-shaped pale spot in front of spinnerets, venter pale gray. Carapace weakly covered with shiny, plumose setae. AME elevated; eye group width 0.84 of caput

width; AME 0.16; ALE 0.12; PME 0.16; PLE 0.12; AME–AME 0.04; AME–ALE 0.02; PME–PME 0.02; PME–PLE 0.04; ALE– PLE 0.02; eye group AME–PME 0.36; AME–AME 0.36; PME–PME 0.34. Clypeus 0.10 high. Abdomen covered with shiny, plumose setae; ALS 0.56 of abdominal length, contiguous. Palp (figs. 81–83): conductor originating medially, with ventrally folded distal end, retrolaterally directed tip,

squared protuberance on base; median apophysis short, ventrally excavated, with two distal tips; sperm duct weakly u-shaped; embolus long, straight, with s-shaped tip; retrolateral tibial apophysis long, triangular, dorsally directed, with distally bent tip.

FEMALE: Total length 3.38. Carapace 1.28 long, 1.24 wide, 0.46 high, length/width 1.03; sternum 0.90 long, 0.80 wide, length/ width 1.12; abdomen 2.10 long, 1.04 wide; coxa I 0.52 long; relative length of coxae I–IV 1.00:0.77:0.73:0.92. Coloration as in male but without dorsal scutum. Eye group width 0.83 of caput width; AME 0.14; PME 0.14; AME– AME 0.02; PME–PLE 0.02; eye group AME–PME 0.32; AME–AME 0.30; PME– PME 0.30. ALS 0.50 of abdominal length. Epigynum (figs. 84, 85): atrium broadly oval, with arched posterior margin; epigynal ducts extending for full width of atrium, in double s-shaped position with spermathecae, spermathecae about their diameter apart, globular, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: New South Wales: near gravel quarry, Bundjalung National park , 29 ° 179S, 153 ° 169E, Feb. 4–Apr. 9, 1963, pitfall (M. Gray, G. Cassis, AMS KS43237 ), 1♀ ; Devils Pulpit State Forest , 29 ° 169S, 153 ° 129E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73952 ), 2♀ ; Doubleduke State Forest , 29 ° 089S, 153 ° 119E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73948 ), 1♀ ; Doubleduke State Forest , 29 ° 129S, 153 ° 169E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73956 ), 1 Oi ; Giberagee State Forest , 29 ° 209S, 153 ° 069E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73950 ), 1♀ ; Giberagee State Forest , 29 ° 229S, 153 ° 059E, Feb. 1997, eucalypt forest pitfall (A. York, AMS KS73951 ), 1♀ ; Jackybulbin Flat Road, 0.3 km S junction Range Road, Doubleduke State Forest , 29 ° 149S, 153 ° 099E, Feb. 4–Apr. 9, 1963, pitfall, elev. 10 m (M. Gray, G. Cassis, AMS KS43240 ), 1♀ ; MacFaydens Road, 2 km WSW junction of Pacific Highway and Glencoe Road , Doubleduke State Forest , 29 ° 129S, 153 ° 149E, Feb. 4–Apr. 9, 1963, pitfall (M. Gray, G. Cassis, AMS KS43238 ), 1♀ ; Morono State Forest , 29 ° 199S, 153 ° 149E, Feb. 1997, eucalypt/paperbark forest pitfall (A. York, AMS KS73949 ), 1 Oi ; Range Road, 1.5 km W junction Lockleys Road, Doubleduke State Forest , 29 ° 149S, 153 ° 109E, Feb. 4–Apr. 9, 1963, pitfall (M. Gray, G. Cassis, AMS KS43241 ), 1 Oi, 2♀ ; junction, Range and Lockleys Roads, Doubleduke State Forest , 29 ° 149S, 153 ° 119E, Feb. 4– Apr. 9, 1963, pitfall (M. Gray, G. Cassis, AMS KS43239 ), 1♀ .

DISTRIBUTION: Known only from northern New South Wales (map 6).