Micronecta annae Kirkaldy, 1905

Tinerella, Paul P., 2013, <strong> Taxonomic revision and systematics of continental Australian pygmy water boatmen (Hemiptera: Heteroptera: Corixoidea: Micronectidae) </ strong>, Zootaxa 3623 (1), pp. 1-121 : 71-78

Micronecta annae Kirkaldy, 1905

( Figs. 41–45 )

Micronecta annae Kirkaldy, 1905: 262 .

Micronecta erato Kirkaldy, 1905: 263 . [synonymy Wróblewski, 1970: 682.]

Micronecta annae: Kirkaldy, 1908: 788 . [list]

Micronecta annae var. pallida: Kirkaldy, 1908: 788 . [synonymy Wróblewski, 1970: 682.]

Micronecta annae: Hale, 1922: 323 .

Micronecta batilla Hale, 1922: 323 . [synonymy Wróblewski, 1970: 682.]

Micronecta annae: Lundblad, 1933: 76 . [list]

Micronecta batilla: Lundblad, 1933: 76 . [list]

Micronecta erato: Lundblad, 1933: 76 . [list]

Micronecta annae: Chen, 1965: 149 . [faunistics]

Micronecta annae: Wróblewski, 1970: 682 . [redescription]

Micronecta annae illiesi Wróblewski, 1970: 687 . [NEW SYNONYMY]

Micronecta annae: Wróblewski, 1972b: 524 . [list]

Micronecta annae illiesi: Wróblewski, 1972b: 524 . [list]

Micronecta batilla: King, 1976 : [stridulation]

Micronecta annae: Wróblewski, 1977: 684 . [faunistics]

Micronecta annae illiesi: Wróblewski, 1977: 684 . [description; distribution]

Micronecta batilla: Bailey, 1983: 35 . [stridulation]

Micronecta annae: Cassis and Gross, 1995: 66 . [catalog]

Micronecta annae illiesi: Cassis and Gross, 1995: 67 . [catalog]

Micronecta annae: Lansbury, 1995: 184 . [faunistics]

Micronecta dixonia King, 1997: 146 . [NEW SYNONYMY]

Micronecta concordia King, 1997: 149 . [NEW SYNONYMY]

Micronecta illiesi: King, 1997: 151 . [species status]

Micronecta annae: Halse et al., 2000: 260 . [list]

Micronecta annae illiesi: Lansbury and Lake, 2002: 3 , 32. [key]

Micronecta annae: Andersen and Weir, 2004: 250 . [key; list]

Micronecta illiesi: Andersen and Weir, 2004: 250 . [key; list]

Diagnosis: This highly variable species is distinguished from all other Australasian Micronectidae by the form of the male genitalia ( Figs. 45d–h ). The combination of the prothoracic lobe ( Fig. 44a ), metaxyphus ( Fig. 44b ), and general facies will permit separation of females. Hemelytral coloration varies from light brown with distinct markings, to dark brown with reduced markings ( Figs. 41c–e , 42a–b , 43a–d ). The darker form of the species seems more prominent in south and eastern Australia (South Australia, Victoria and Tasmania).

Size: ( Table 14). Brachypterous form: 2.74–3.44. Macropterous form: 3.18–3.62. Complete measurement data for 20 specimens of M. annae are included in Table 14.

Derivation of specific epithet: Kirkaldy probably named the species for his wife, Annie K. Brenham (see Dolling 1991 for short biography of Kirkaldy).

Notes on type material: The whereabouts of the type material are unknown. Kirkaldy noted in the original description (1905: 263) “my collection”, indicating he retained the type ( Dolling 1991). The type was not located in BPBM or SEMC holdings.

I was able to examine Hale’s (1922) types of Micronecta batilla from SAMA, confirming Wróblewski’s synonymy with M. annae . Owing to the inability to locate Kirkaldy’s holotype of M. annae , I have illustrated Hale’s holotype of M. batilla for comparative purposes. Holotype (♂) and allotype (♀) on point card ( Fig. 41a–c ): “Macclesfield [Mt. Lofty Ranges], South Australia. H. M. Hale”. Pin bearing small handwritten label: “Type” and large handwritten label ( Fig. 41b ): “ Micronecta batilla Hale , South Australia.” The number “I15187” handwritten in upper right-hand corner of label. “Type” handwritten in red ink, along right side of label. On a separate pin, the paratype series consists of 16 specimens glued on a rectangular point card and labeled as “Co-Type” on pink label, same data as above; larger label beneath, handwritten and bearing “ Micronecta batilla Hale ,” “ Paratype ” printed in red along right margin of label.

At present, I decline designating a neotype of M. annae , as all possible locations of Kirkaldy’s type have not been exhausted. Moreover, the present definition of M. annae seems quite clear collectively from Kirkaldy’s (1905) description, Wróblewski’s (1970) subsequent redescription and synonymy of M. batilla , and further study of specimens from this research. In the interim, Hale’s type of M. batilla = M. annae , which agrees well with Kirkaldy’s original description, serves as proxy for the concept of M. annae .

Description: Based on brachypterous and macropterous forms, differences noted. Measurements. Length: male 2.98–3.48; female 2.87–3.62; Width: male 1.63–1.77; female 1.61–1.84; Width of head: male 1.09–1.25; female 1.04–1.27; Synthlipsis: male 0.48–0.53; female 0.48–0.54; Width of eye: male 0.25–0.36; female 0.21–0.36; Width of pronotum: male 1.16–1.29; female 1.10–1.31; Length of pronotum: male 0.22–0.33; female 0.21–0.34.

Color: Ground color brown ( Figs. 41c–d , 42a–b , 43a–d ). Head light brown, eyes reddish-brown to silveryblack. Vertex light brown with central reddish longitudinal line extending onto light brown colored frons. Clypeus and genae dark brown, rastrate, with evenly distributed small pitting. Labium black. Antennae light brown, same color as vertex and frons. Pronotum in macropterous form unicolorous light brown, with darker areas restricted to lateral portions. Pronotum in brachypters light brown with dark area present as a transverse band spanning entire pronotum. Scutellum pale yellowish-brown. Clavus dark brown with basal pale, diagonal area prominent in macropterous form. Darker coloration of clavus in macropterous form restricted laterally, often as a narrow, dark longitudinal line. Clavus in brachypterous forms generally lighter brown with prominent dark longitudinal areas laterally. Corium of both forms with variably developed longitudinal dark lines, ranging from bold, distinct patterning over entire corium to restricted, interrupted lines laterally. Prenodal embolar area long and infuscated, well-developed in both wing forms. Postnodal embolar widened and short in both wing forms, infuscation prominent in macropterous specimens examined. Left membrane in both sexes subhyaline, often dark brown to black in macropters; inner margin light brown to pale whitish. Venter of both sexes dark brown to black, natatorial setae of metatarsus embrowned.

Structural characteristics: Ratio of body length/width: males 1.91; females 1.90 Metrics of macropters proportionately larger than brachypters. Head width nearly equal to pronotal width, synthlipsis 1.7 times as wide as posterior width of eye. Ocular index: males 1.50; females 1.62. General facies of head (vertex, frons, and labium) proportionate. Third antennal segment densely pilose, short and broad, upper margin straight, lower margin very broadly rounded, apex acuminate, segments one and two very short. Pronotum robust in macropters, convex and widest at middle with lateral margins tapered, about four times as wide as long (W/L: males 1.29/0.32; females 1.31/0.34). Pronotum in brachypterous form proportionately shorter, lateral margins tapered, nearly pointed, averaging 4.75 times as wide as long (W/L: males 1.14/0.24; females 1.10/0.21). Both wing forms with developed anterior carina. Prothoracic lobe short and comparatively wide, with lower and posterior lobes broadly rounded, anterior margin acutely pointed ( Fig. 44a ). Hemelytral microsculpturing of both wing morphs consisting of very fine transverse meshes. General hemelytral facies of macropterous form rastrate, that of brachypter smooth. Longer, spinelike setae uniformly distributed and widely spaced over entire hemelytral surface. Metathoracic wings of macropter well-developed, reaching nearly to apices of hemelytra. Metathoracic wings of brachypterous form atrophied, very short, and reaching to abdominal tergite V. Lateral spines on abdominal segments IV–VIII: IV: one stout, short spine, two thin, long setae; V: two stout, short setae, one long, thin seta; VI: one stout, short seta, one long, thin seta; VII: three short, stout spines, one long, stout spine, one long, thin seta; VIII: six stout, short spines, three long, thin setae. Metaxyphus of both sexes narrowed, triangular, with apex sharply pointed ( Figs. 44b , 45a ).

Male foreleg ( Fig. 44c ): femur setose basally, with two short, stout spines in basal third near ventral surface. Tibia with one thin seta basally and one longer, stout spine in apico-ventral portion. Pala with six longer setae dorsally; palmar area with nine setae in upper row, 12–14 setae in lower row, lower row setae more pronounced than those of dorsal row. Apex of pala with a long thickened seta. Palar claw ( Figs. 44d , 45b ) robust, spatulate. Female foreleg with same general setal arrangement as male. Mesotarsal claws long, about one-half the length of the mesotarsus.

Lateral lobes of abdominal tergum IV broadly rounded. Right lobe slightly longer than left, with 17 long setae, left lobe shorter, with 14 long evenly spaced, apically placed setae. Prestrigilar flap of tergal segment V as in Figure 44e . Strigil well-developed, ovate, with few combs. Median lobe of sternite VII ( Fig. 44f ) long, with numerous setae restricted throughout, apex narrow and rounded. Free lobe of tergite VIII ( Figs. 44g , 45c ) broad, outer angle broadly rounded with short, well-produced inner angle. Inner angle with about 27 long setae, outer upper angle with 7–12 short setae. Pars stridens processus cleaner ridges of tergite VIII not examined. Male left paramere ( Figs. 45d–f ) relatively long, shaft convexly curved, gradually widening distally, area immediately before apex expanded, apical portion of paramere with short, acute hook ( Figs. 45d–e ). Scales present in apical third of paramere, base broad, lower portion broadly rounded ( Figs. 45d–f ). Right paramere ( Figs. 45d–g ) long, parallelsided, with tip tapering to blunt point ( Figs. 45d–g ). Base widened, lower portion lobate, with about 39 plectral ribs confined to lower portion ( Fig. 45h ). Aedeagus broad, as in Figures 45c–e .

Distribution and Habitat: AUSTRALIA: [widespread]: Australian Capital Territory, New South Wales, Queensland, South Australia, Tasmania, Victoria, Western Australia. A very common species in Australia, collected in a variety of aquatic habitats, as well as commonly taken at lights. So far, not known from Northern Territory.

Discussion: Micronecta annae is a widespread, highly variable species occurring throughout continental Australia. Much of the confusion surrounding the species and defining the variability seen, is owed to Kirkaldy’s short, imprecise original description and the whereabouts of the type specimen. Further, neither Hale (1922) or Chen (1965) treated the species, only repeating Kirkaldy’s original description. At that time, Hale described M. batilla (= M. annae ) and made no additional comments on the identity of M. annae . In her investigation, Chen (1965) recognized Hale’s (1922) M. batilla and commented that it was impossible to recognize M. annae based on Kirkaldy’s description.

Wróblewski (1970) was the first to redescribe M. annae , based on examination of Kirkaldy’s description, study of Hale’s M. batilla , and additional specimens of M. annae . He provided detailed figures of the species, synonymized M. batilla and M. erato with M. annae , and provided brief comments on M. annae var. pallida (= M. annae ). At the same time, he described M. annae illiesi from the limited material he had available (2♂, 17♀♀), noting the similarities of his new subspecies with M. annae . Wróblewski (1977) later amended his original description based on a much larger collection of material from Victoria and Tasmania. There he noted the original metrics for M. illiesi were less divergent than first reported. I was able to examine most of the material the redescription was based on ( Wróblewski 1977: 684) and have identified all of it as M. annae .

I have examined all characters discussed by Wróblewski (1970, 1972b) as diagnostic for M. annae illiesi and regard these as within the variation of the species. These characters include hemelytral patterning, which is highly variable (see above), the synthlipsis (interocular space vs. eye width), which varies greatly within and among short and long winged forms ( Table 14), and the metaxyphus, sternal process of abdominal segment VII, and free lobe abdominal tergite VIII, which are variable in numerous samples examined from across Australia. Further, microscopy examination of the male genitalia (both light and scanning electron microscopy) reveals that rotation of the pygophore produces paramere morphology seen in illustrations for both M. annae and M. illiesi ( Figs. 45d–h ).

Based on my morphology examination of over 6,000 specimens of M. annae and more than 3,500 specimens of material previously determined as M. illiesi , I herein propose synonymy of M. annae . and M. illiesi . All differences described previously with morphology ( Wróblewski 1970, 1977) are regarded as natural variation within the species. Further study using molecular markers (e.g., mitochondrial cytochrome oxidase I) across populations will be useful for analyzing any evidence of divergence within M. annae .

Micronecta concordia and M. dixonia are also herein synonymized with M. annae . I examined the type specimens for both of these at MVMA and found these morphologically identical with M. annae . King (1997) based his description of both species nearly entirely on acoustic signals he recorded from (and nearby) each type locality. Sound recordings of both species were made within a restricted geographical area, often only from one large pond, which would seemingly limit any analysis of variation in mating calls and chorusing of Micronecta species discussed by King (1997, 1999a, 1999b, 1999c). Additional character study of bioacoustics and the use of molecular markers and ecology, would prove useful for determination of character states beyond morphology.

Specimens examined: AUSTRALIA: Australian Capital Territory: Canberra. 30-III-1971. R. Wilcox. (2♂, 2♀♀). [ USNM] ; Australian Capital Territory: Acton Dickson Road ( CSIRO). 11-XI-1980. M. Pion & C. Reichart. (1♀). [ USNM] ; New South Wales: Moonbar, Monaco , 3-3500 ft. III-1889. Helms. (1♀). [ AMSA] ; New South Wales: Racecourse. 11-VII-1961. D. J. Woodland. (5♀♀). [ USNM] ; New South Wales: Armidale Dist., Gara River, across Grafton Hwy. 15-VII-1961. D. J. Woodland. (36♂, 39♀♀). [ USNM] ; New South Wales: Commissioner’s Bridge, across Grafton Hwy. 15-VII-1961. D. J. Woodland. (12♂, 32♀♀). [ USNM] ; New South Wales: Pt. Lookout National Park, Serpentine River . 13-VIII-1961. D. J. Woodland. (11♂, 43♀♀). [ USNM] ; New South Wales: Kingston. 25-VIII-1961. D. J. Woodland. (1♂, 1♀). [ USNM] ; New South Wales: Westbury Bridge. 25-VIII-1961. D. J. Woodland. (1♀). [ USNM] ; New South Wales: Guyra-Tingha Road, Brooklet into Limeston Creek . 5-IX-1961. D. J. Woodland. (1♀). [ USNM] ; New South Wales: Junction, Guyra-Tingha Road, Ollera Creek . 5-IX-1961. D. J. Woodland. (3♂, 7♀♀). [ USNM] ; New South Wales: Shell Harbour. 3-VIII-1962. Muddy waterhole, stagnant creek. A. W. Sweeney. (3♂, 5♀♀). [ USNM] ; New South Wales: Ando, McLachlan River . 18-XI-1965. A. Neboiss. (2♂, 2♀♀). [ MVMA] ; New South Wales: Murray River, Wingillre Station . 27- VI-1967. R. Blackwood. (2♂, 10♀♀). [ MVMA] ; New South Wales: Apsley River, Apsley Falls . 31 03 05S; 151 45 59E. 2-X-1994. Edge. E. Turak. (13♂, 11♀♀). [ AMSA] ; New South Wales: Caparra Creek, Caparra Hut. 31 42 07S; 152 11 26E. 4-X-1994. Edge. Joanne Ling. (2♂, 6♀♀). [ AMSA] ; New South Wales: Abercrombie River, Speary Ridge. 34 10 12S; 149 56 08E. 25-X-1994. Edge. Joanne Ling. (2♂, 3♀♀). [ AMSA] ; New South Wales: Billabong Creek, Culcairn Bridge. 35 40 21S; 147 02 03E. 25-X-1994. Edge. Frances Laurenson. (12♂, 10♀♀). [ AMSA] ; New South Wales: Billabong Creek, u/s Jerilderie. 35 21 11S; 145 45 39E. 25-X-1994. Macrophytes. Frances Laurenson. (1♀). [ AMSA] ; New South Wales: Murray River, Circular Log Landing. 35 59 16S; 144 55 16E. 25-X-1994. Edge. E. Turak. (12♂, 9♀♀). [ AMSA] ; New South Wales: Murray River, u/s Moama Bama State Forest . 36 04 58S; 144 50 47E. 25-X-1994. Edge. E. Turak. (2♂, 1♀). [ AMSA] ; New South Wales: Tuena Creek, d/s Bear’s Gully. 34 04 28S; 149 22 08E. 25-X-1994. Edge. Joanne Ling. (4♀♀). [ AMSA] ; New South Wales: Abercrombie River, u/s Isabella River . 34 01 56S; 149 29 48E. 26-X-1994. Edge. Joanne Ling. (1♂, 3♀♀). [ AMSA]; same data. Macrophytes. Joanne Ling. (4♂, 2♀♀). [ AMSA] ; New South Wales: Lachlan River, d/s Humes Creek . 33 44 31S; 149 17 50E. 26-X-1994. Edge. Joanne Ling. (1♂, 4♀♀). [ AMSA] ; New South Wales: Murray River, Howlong Common. 35 59 40S; 146 37 02E. 26-X-1994. Macrophytes. F. Laurenson. (6♂, 9♀♀). [ AMSA] ; New South Wales: Wakool River, Noorang State Forest . 35 19 55S; 143 55 55E. 26-X-1994. Edge. E. Turak. (7♂, 6♀♀). [ AMSA] ; New South Wales: Grabben Gullen Creek, u/s Lachlan River . 34 32 25S; 149 09 27E. 27-X-1994. Macrophytes / Edge. Joanne Ling. (3♂, 14♀♀). [ AMSA] ; New South Wales: Murray River, u/s Indi Bridge. 36 14 15S; 148 02 02E. 27-X-1994. Edge. J. O’Donnell. (1♂). [ AMSA] ; New South Wales: Murrumbidgee River, d/s Maude Weir. 34 28 31S; 144 17 20E. 27-X-1994. Edge. E. Turak. (10♂, 7♀♀). [ AMSA] ; New South Wales: Belubula River, Taplow Flat. 33 35 34S; 148 52 13E. 28-X-1994. Edge. Joanne Ling. (1♂, 6♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Long Beach. 34 39 05S; 146 22 41E. 28-X-1994. E. Turak. (12♂, 10♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Meriola Reserve . 34 28 38S; 145 04 18E. 28-X-1994. Edge. E. Turak. (4♂, 3♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Nulabor. 34 28 32S; 145 06 54E. 28-X-1994. Edge. E. Turak. (8♂, 3♀♀). [ AMSA] ; New South Wales: Lachlan River, Straney’s Bridge. 33 22 30S; 147 39 13E. 29-X-1994. Edge. R. Cowell. (5♂, 5♀♀). [ AMSA] ; New South Wales: Lachlan River, u/s Mandagery Creek . 33 30 07S; 148 18 29E. 29-X-1994. Edge. R. Cowell. (2♂, 5♀♀). [ AMSA] ; New South Wales: Lachlan River, Murrin Bridge. 33 12 18S; 146 21 38E. 30-X-1994. Edge. R. Cowell. (1♂). [ AMSA] ; New South Wales: Mann River, at Mann River Falls , CLAR 01. 29 42 56S; 151 56 24E. 31-X-1994. Edge. Eren Turak. (4♂, 6♀♀). [ AMSA] ; New South Wales: Towamba River, Burragate. 36 59 37S; 149 36 29E. 15-III-1995. Edge. M. Eggerup. (1♀). [ AMSA] ; New South Wales: Abercrombie River, Speary Ridge. 34 10 12S; 149 56 08E. 23-III-1995. Macrophytes. Russell Cowell. (3♂, 9♀♀). [ AMSA] ; New South Wales: Belubula River, Taplow Flat. 33 35 34S; 148 52 13E. 23-III-1995. Edge. R. Cowell. (1♂, 2♀♀). [ AMSA]; same data: B. Gardiner. (2♂, 1♀). [ AMSA] ; New South Wales: Rocky Bridge Creek, u/s Piesley’s Creek . 33 51 33S; 149 08 14E. 24-III-1995. Edge. Russell Cowell. (3♂, 3♀♀). [ AMSA] ; New South Wales: Abercrombie River, Speary Ridge. 34 10 12S; 149 56 08E. 25-III-1995. Edge. Frances Laurenson. (2♂, 4♀♀). [ AMSA] ; New South Wales: Abercrombie River, u/s Isabella River . 34 01 56S; 149 29 48E. 25-III-1995. Edge. F. Laurenson. (6♂, 4♀♀). [ AMSA] ; New South Wales: Box Creek Esc. Chan., Conargo Road. 35 24 11S; 145 03 27E. 25-III-1995. Edge. B. Gardiner. (1♀). [ AMSA] ; New South Wales: Murray River, u/s Indi Bridge. 36 14 15S; 148 02 02E. 25-III-1995. Edge. E. Turak. (2♂, 2♀♀). [ AMSA] ; New South Wales: Boorowa River, at Prosser’s Crossing. 34 08 38S; 148 48 33E. 26-III- 1995. Macrophytes. F. Laurenson. (1♀). [ AMSA] ; New South Wales: Grabben Gullen Creek, u/s Lachlan River . 26-III-1995. Edge. R. Cowell. (1♂, 2♀♀). [ AMSA] ; New South Wales: Lachlan River, d/s Humes Creek . 33 44 31S; 149 17 50E. 26-III-1995. Macrophytes. R. Cowell. (4♂, 6♀♀). [ AMSA] ; New South Wales: Lachlan River, Narrawa Mtn. 34 22 28S; 149 06 36E. 26-III-1995. Edge. Frances Laurenson. (9♂, 8♀♀). [ AMSA] ; New South Wales: Murray River, Jingellic. 35 55 54S; 147 42 06E. 26-III-1995. Edge. E. Turak. (14♂, 6♀♀). [ AMSA] ; New South Wales: Murray River, Nursery Bend #2. 35 46 01S; 149 20 51E. 26-III-1995. Edge. B. Gardiner. (5♂, 3♀♀). [ AMSA] ; New South Wales: Niemur River, Nacurrie Road. 35 14 04S; 143 58 11E. 26-III-1995. Edge. Bruce Gardiner. (8♂, 7♀♀). [ AMSA] ; New South Wales: Wakool River, Noorang State Forest . 35 19 55S; 143 55 55E. 26-III-1995. Edge. Joanne Ling. (2♀♀). [ AMSA] ; New South Wales: Billabong Creek, u/s Jerilderie. 35 21 11S; 145 45 39E. 27-III-1995. Macrophytes. E. Turak. (4♂, 9♀♀). [ AMSA] ; New South Wales: Lachlan River, Murrin Bridge. 33 12 18S; 146 21 38E. 27-III-1995. Edge. R. Cowell. (1♀). [ AMSA] ; New South Wales: Lachlan River, Straney’s Bridge. 33 22 30S; 147 39 13E. 27-III-1995. Edge. Russell Cowell. (5♂, 14♀♀). [ AMSA] ; New South Wales: Lachlan River, u/s Mandagery Creek . 33 30 07S; 148 18 29E. 27-III-1995. Edge. F. Laurenson. (12♂, 6♀♀). [ AMSA] ; New South Wales: Murray River, Barham Mill Bend. 35 44 32S; 144 12 54E. 27-III-1995. Edge. Joanne Ling. (15♂, 11♀♀). [ AMSA] ; New South Wales: Murray River, Howlong Common. 35 59 40S; 146 37 02E. 27-III-1995. Edge. E. Turak. (11♂, 8♀♀). [ AMSA] ; New South Wales: Murray River, Howlong Common. 35 59 40S; 146 37 02E. 27-III-1995. Macrophytes. E. Turak. (8♂, 12♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, d/s Maude Weir. 34 28 31S; 144 17 20E. 27-III-1995. Edge. D. Jarvis. (16♂, 8♀♀). [ AMSA] ; New South Wales: Wakool River, u/s Tarara Crossing. 34 55 29S; 143 28 39E. 27-III-1995. Edge. Joanne Ling. (5♂, 5♀♀). [ AMSA] ; New South Wales: Billabong Creek, d/s Coleambally, Outfall Dam. 35 02 44S; 144 26 48E. 28-III-1995. Edge. D. Jarvis. (8♂, 10♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Long Beach. 34 39 05S; 146 22 41E. 28-III-1995. Edge. F. Laurenson. (16♂, 12♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Nulabor. 34 28 32S; 145 06 54E. 28-III-1995. Edge. Frances Laurenson. (8♂, 9♀♀). [ AMSA] ; New South Wales: Holey Flat Creek, u/s Moorland. 31 45 39S; 152 38 12E. 6-IV-1995. Edge. E. Turak. (1♀). [ AMSA] ; New South Wales: Manning River, Henry’s Bridge. 31 52 55S; 151 29 21E. 8-IV-1995. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Barrington River, Bob’s Crossing. 32 01 45S; 151 27 60E. 10-IV-1995. Edge. Joanne Ling. (1♀). [ AMSA] ; New South Wales: Peacock Creek . 28 44 48S; 152 37 04E. 9-V- 1995. Edge. E. Turak. (7♂, 11♀♀). [ AMSA] ; New South Wales: Cataract River, at Sandy Hill. 28 55 33S; 152 13 31E. 10-V-1995. Macrophytes. E. Turak. (1♀). [ AMSA] ; New South Wales: Chander’s Creek, Morera State Forest . 29 55 27S; 152 31 05E. 10-V-1995. Edge. Joanne Ling. (2♂). [ AMSA] ; New South Wales: Corall’s Creek, at Bald Rock National Park . 28 50 51S; 152 05 54E. 10-V-1995. Edge. E. Turak. (19♂, 28♀♀). [ AMSA] ; New South Wales: Mann River Falls . 11-V-1995. Edge. E. Turak. (11♂, 19♀♀). [ AMSA] ; New South Wales: Washpool Creek, Washpool National Park . 29 16 20S; 152 25 47E. 13-V-1995. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Nepean River, u/s Penrith. 33 48 12S; 150 37 36E. 26-V-1995. Edge. Frances Laurenson. (2♂, 8♀♀). [ AMSA] ; New South Wales: Clarence River, at Carnham Crossing. 29 20 09S; 152 32 07E. 9-X-1995. Edge. E. Turak. (2♀♀). [ AMSA] ; New South Wales: Wash Pool Creek, Wash Pool National Park , CLAR 07. 29 16 20S; 152 25 47E. 9-X-1995. Riffle. Alexander Leask. (1♂). [ AMSA] ; New South Wales: Mann River Falls . 29 42 56S; 151 56 20E. 11-X-1995. Riffle. Joanne Ling. (1♂). [ AMSA] ; New South Wales: EPA Survey MRHI, Allyn River at Halton , HUNT 03. 32 18 27S; 151 30 39E. 18-X-1995. Edge. Alexander Leask. (2♂, 2♀♀). [ AMSA] ; New South Wales: Murray River, Circular Log Landing. 35 59 16S; 144 55 16E. 25-X-1995. Edge. B. Gardiner. (15♂, 9♀♀). [ AMSA] ; New South Wales: Murray River, u/s Moama Bama State Forest . 36 04 58S; 144 50 47E. 25-X-1995. Edge. B. Gardiner. (14♂, 12♀♀). [ AMSA] ; New South Wales: Nepean River, d/s Penrith, BMG Causeway. 33 44 06S; 150 40 38E. 31-X-1995. Edge. Joanne Ling. (3♂, 10♀♀). [ AMSA] ; New South Wales: Nepean River, u/s Penrith. 33 48 12S; 150 37 36E. 31-X-1995. Edge. Joanne Ling. (1♂, 3♀♀). [ AMSA] ; New South Wales: Lachlan River, d/s Humes Creek . 33 44 31S; 149 17 50E. 6-XI-1995. Macrophytes. R. Cowell. (2♀♀). [ AMSA] ; New South Wales: Billabong Creek, d/s Coleambally, Outfall Dam. 35 02 44S; 144 26 48E. 7- XI-1995. Edge. Joanne Ling. (3♂, 10♀♀). [ AMSA] ; New South Wales: Murray River, Barham Mill Bend. 35 44 32S; 144 12 54E. 7-XI-1995. Edge. Joanne Ling. (14♂, 9♀♀). [ AMSA] ; New South Wales: Murray River, Windomal Landing. 34 47 19S; 149 19 49E. 7-XI-1995. Edge. Joanne Ling. (8♂, 3♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, d/s Maude Weir. 34 28 31S; 144 17 20E. 7-XI-1995. Edge. Joanne Ling. (1♂, 1♀). [ AMSA] ; New South Wales: Murrumbidgee River, Long Beach. 34 39 05S; 146 22 41E. 7-XI-1995. Edge. E. Turak. (9♂, 13♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, Meriola Reserve . 34 28 38S; 145 04 18E. 7-XI-1995. Edge. E. Turak. (6♂, 6♀♀). [ AMSA] ; New South Wales: Niemur River, Nacurrie Road. 35 14 04S; 143 58 11E. 7-XI-1995. Edge. Natacha Waddell. (4♂, 2♂). [ AMSA] ; New South Wales: Wakool River, Noorang State Forest . 35 19 55S; 143 55 55E. 7-XI-1995. Edge. B. Gardiner. (2♂, 2♀♀). [ AMSA] ; New South Wales: Wakool River, u/s Tarara Crossing. 34 55 29S; 143 28 39E. 7-XI-1995. Edge. Joanne Ling. (9♂, 7♀♀). [ AMSA] ; New South Wales: Lachlan River, u/s Mandagery Creek . 33 30 07S; 148 18 29E. 8-XI-1995. Edge. E. Turak. (4♂, 3♀♀). [ AMSA] ; New South Wales: Murray River, Circular Log Landing. 35 59 16S; 144 55 16E. 8- XI-1995. Edge. Joanne Ling. (5♂, 2♀♀). [ AMSA] ; New South Wales: Murray River, u/s Moama Bama State Forest . 36 04 58S; 144 50 47E. 8-XI-1995. Edge. Natacha Waddell. (16♂, 2♀♀). [ AMSA] ; New South Wales: Belubula River, Taplow Flat. 33 35 34S; 148 52 13E. 9-XI-1995. Edge. Joanne Ling. (4♂, 2♀♀). [ AMSA] ; New South Wales: Murray River, Howlong Common. 35 59 40S; 146 37 02E. 9-XI-1995. Edge. Joanne Ling. (6♂, 11♀♀). [ AMSA] ; New South Wales: Billabong Creek, u/s Jerilderie. 35 21 11S; 145 45 39E. 10-XI-1995. Edge. Natacha Waddell. (8♂, 17♀♀). [ AMSA] ; New South Wales: Box Creek Esc. Chan., Conargo Road. 35 24 11S; 145 03 27E. 10-XI-1995. Edge. Joanne Ling. (10♂, 12♀♀). [ AMSA] ; New South Wales: Grabben Gullen Creek, u/s Lachlan River . 34 32 25S; 149 09 27E. 15-XI-1995. Edge. Alexander Leask. (1♂, 2♀♀). [ AMSA]; same data: Macrophytes. R. Cowell. (2♂, 7♀♀). [ AMSA] ; New South Wales: Murray River, Jingelic. 35 55 54S; 147 42 06E. 22-XI-1995. Edge. N. Meucci. (8♂, 12♀♀). [ AMSA] ; New South Wales: Eucumbene River, Gouge Station . 25-XI-1995. Edge. Joanne Ling. (2♂, 1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Shoalhaven River, Fossicker’s Flat , SHOR 05. 34 49 16S; 150 49 16E. 5-XII-1995. Edge. Joanne Ling. (36♂, 18♀♀). [ AMSA] ; New South Wales: Shoalhaven River, Fossicker’s Flat. 34 49 16S; 150 11 32E. 5-XII-1995. Edge. Joanne Ling. (3♂, 3♀♀). [ AMSA] ; New South Wales: Colo River, near Moran’s Rock. 33 37 50S; 151 14 26E. 8-XII-1995. Edge. E. Turak. (1♀). [ AMSA] ; New South Wales: Shoalhaven River, d/s Tallowa Dam. 34 46 50S; 150 18 54E. 12-XII-1995. Edge. Joanne Ling. (1♂, 1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Goulburn River near Turnbridge Creek , HUNT 07. 32 18 37S; 150 14 19E. 13-XII-1995. Edge. Natacha Waddell. (120♂, 98♀♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Kiernam’s Creek off Dart Brook , HUNT 08. 31 52 37S; 150 39 49E. 13-XII-1995. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Grabben Gullen Creek, u/s Lachlan River . 34 32 25S; 149 09 27E. 18-III-1996. Edge. Joanne Ling. (10♂, 12♀♀). [ AMSA]; same data: Macrophytes. Joanne Ling. (13♂, 7♀♀). [ AMSA] ; New South Wales: Lachlan River, d/s Humes Creek . 33 44 31S; 149 17 50E. 18-III-1996. Edge. Joanne Ling. (9♂, 17♀♀). [ AMSA] ; New South Wales: Lachlan River, d/s Kiacatoo Bridge. 33 03 14S; 146 46 05E. 19-III-1996. Edge. Joanne Ling. (1♀). [ AMSA] ; New South Wales: Lachlan River, Straney’s Bridge. 33 22 30S; 147 39 13E. 19-III-1996. Edge. Joanne Ling. (3♂, 5♀♀). [ AMSA] ; New South Wales: Lachlan River, u/s Cowra. 33 53 17S; 148 45 03E. 19-III-1996. Edge. Joanne Ling. (6♂, 17♀♀). [ AMSA] ; New South Wales: Murray River, d/s W Albury Sewage Plant. 36 04 27S; 146 52 12E. 19-III-1996. Edge. Natacha Waddell. (13♂, 15♀♀). [ AMSA] ; New South Wales: Stony Creek . 35 39 26S; 147 45 32E. 19-III- 1996. Edge. Natacha Waddell. (2♀♀). [ AMSA] ; New South Wales: Bago Creek . 35 33 57S; 148 01 58E. 20-III- 1996. Edge. Natacha Waddell. (2♂). [ AMSA] ; New South Wales: Billabong Creek, Culcairn Bridge. 35 40 21S; 147 02 03E. 20-III-1996. Edge. Natacha Waddell. (15♂, 17♀♀). [ AMSA] ; New South Wales: Murray River, Windomal Landing. 34 47 19S; 149 19 49E. 20-III-1996. Edge. Joanne Ling. (2♂, 7♀♀). [ AMSA] ; New South Wales: Murrumbidgee River, d/s Maude Weir. 34 28 31S; 144 17 20E. 20-III-1996. Edge. M. Root. (1♂, 1♀). [ AMSA] ; New South Wales: Wakool River, Noorang State Forest . 35 19 55S; 143 55 55E. 20-III-1996. Edge. M. Root. (1♂, 1♀). [ AMSA] ; New South Wales: Boorowa River, at Prosser’s Crossing. 34 08 38S; 148 48 33E. 21-III- 1996. Edge. A. McLean. (7♂, 4♀♀). [ AMSA] ; New South Wales: Box Creek Esc. Chan., Conargo Road. 35 24 11S; 145 03 27E. 21-III-1996. Edge. M. Root. (14♂, 9♀♀). [ AMSA] ; New South Wales: Murray River, Honeymoon Bend. 35 49 52S; 145 09 28E. 21-III-1996. Edge. Murray Root. (10♂, 9♀♀). [ AMSA] ; New South Wales: Abercrombie River, Speary Ridge. 34 10 12S; 149 56 08E. 22-III-1996. Edge. Adam McLean. (1♂, 3♀♀). [ AMSA] ; New South Wales: Murray River, Circular Log Landing. 35 59 16S; 144 55 16E. 22-III-1996. Edge. Murray Root. (5♂, 2♀♀). [ AMSA] ; New South Wales: Murray River, u/s Moama Bama State Forest . 36 04 58S; 144 50 47E. 22-III-1996. Edge. Murray Root. (15♂, 13♀♀). [ AMSA] ; New South Wales: Clarence River, at Carnham Crossing , CLAR 17. 29 20 09S; 152 32 07E. 14-IV-1996. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Nymboida River, Dalmorton SF/ Nymboida NP. 29 42 38S; 152 33 31E. 14-IV-1996. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Wash Pool Creek, Wash Pool National Park , CLAR 07. 29 16 20S; 152 25 47E. 14-IV-1996. Edge. Natacha Waddell. (3♂). [ AMSA] ; New South Wales: Little Dandahra Creek, Mulligan’s Hut , CLAR 14. 29 31 02S; 152 21 33E. 14-IV-1996. Edge. Joanne Ling. (1♂). [ AMSA] ; New South Wales: Clarence River, Paddy’s Flat , CLAR 16. 28 43 25S; 152 24 21E. 17-IV-1996. Edge. Joanne Ling. (6♂, 3♀♀). [ AMSA] ; New South Wales: Chandler’s Creek, Morera State Forest , CLAR 09. 29 55 27S; 152 31 05E. 19-IV-1996. Edge. Joanne Ling. (2♂, 2♀♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Little Nymboida, Confluens Nymboidea R., CLAR 20. 30 04 16S; 152 45 18E. 19-IV-1996. Edge. Joanne Ling. (6♂, 5♀♀). [ AMSA] ; New South Wales: Golden Moon Creek, up Little River . 34 09 34S; 150 26 32E. 14-V-1996. Edge. Joanne Ling. (3♀♀). [ AMSA] ; New South Wales: Nepean River, d/s Penrith, BMG Causeway. 33 44 06S; 150 40 38E. 14-V-1996. Macrophytes. Natacha Waddell. (10♂, 19♀♀). [ AMSA] ; New South Wales: Shoalhaven River, Fossicker’s Flat. 34 49 16S; 150 11 32E. 15-V-1996. Edge. E. Turak. (9♂, 23♀♀). [ AMSA] ; New South Wales: Shoalhaven River, d/s Tallowa Dam. 34 46 50S; 150 18 54E. 16-V-1996. Edge. Joanne Ling. (1♂, 3♀♀). [ AMSA] ; New South Wales: Kangaroo River, up Gerrigong Creek . 34 41 17S; 150 35 58E. 18-V-1996. Edge. Joanne Ling. (2♂, 1♀). [ AMSA] ; New South Wales: Jocks Creek, u/s Cox’s River . 33 34 18S; 150 04 38E. 24-V-1996. Edge. E. Turak. (2♀♀). [ AMSA] ; New South Wales: Nepean River, u/s Penrith. 33 48 12S; 150 37 36E. 29-V-1996. Edge. Eren Turak. (2♂, 1♀). [ AMSA] ; New South Wales: Murrumbidgee River, d/s Tantagara Reservoir. 35 48 02S; 148 40 32E. 17-III-1997. Edge. A. Leask. (1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Murray River - Tom Groggin , MURR 24. 36 32 52S; 148 07 34E. 21-III-1997. Edge. Alexander Leask. (2♂, 3♀♀). [ AMSA] ; New South Wales: Cooma Creek, u/s Mittagong Road. 36 11 37S; 149 07 01E. 23-III-1997. Edge. A. Leask. (4♂, 3♀♀). [ AMSA] ; New South Wales: Snowy River, near Pats Patch. 36 26 43S; 148 38 21E. 23-III-1997. Edge. T. Kobayashi. (6♂, 5♀♀). [ AMSA] ; New South Wales: Bombala River at Cambalong. 36 53 25S; 149 06 41E. 24- III-1997. Edge. A. Leask. (5♂, 7♀♀). [ AMSA] ; New South Wales: Areluen Creek, at Neringla Road. 35 39 33S; 149 48 21E. 25-III-1997. Edge. Natacha Waddell. (1♀). [ AMSA] ; New South Wales: Gillamatoney Creek, d/s Braidwood , SHOA 501. 35 26 37S; 149 46 45E. 25-III-1997. Edge. Natacha Waddell. (1♂). [ AMSA] ; New South Wales: Major’s Creek, at Major’s Creek Road. 35 33 31S; 149 45 01E. 25-III-1997. Edge. Natacha Waddell. (2♂, 1♀). [ AMSA]; same data: Macrophytes. Natacha Waddell. (4♂, 1♀). [ AMSA] ; New South Wales: Kangaroo River, d/s Hampton Bridge , SHOA 503. 34 43 43S; 150 31 11E. 26-III-1997. Edge. Alexander Leask. (2♂, 3♀♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Oaky Creek, at Richmond Range SF. 28 37 41S; 152 45 49E. 7-V-1997. Edge. Eren Turak. (4♂, 1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Lynch’s Creek, Rich 515. 28 25 40S; 153 01 32E. 9-V-1997. Edge. Graham Sherwin. (1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Roseberry Creek, Rich 505. 28 28 29S; 152 49 24E. 9-V-1997. Edge. Alexander Leask. (1♂, 1♀). [ AMSA] ; New South Wales: EPA Survey MRHI, Rocky Creek, Rich 01. 28 34 18S; 153 20 33E. 11-V-1997. Edge. Alexander Leask. (3♂). [ AMSA] ; New South Wales: Shoalhaven River, Fossicker’s Flat , SHOA 05. 34 49 16S; 150 11 32E. 11-VI-1997. Edge. Alexander Leask. (11♂, 5♀♀). [ AMSA] ; New South Wales: Kangaroo River, up Gerrigong Creek , SHOA 01. 34 41 17S; 150 35 58E. 26-XI-1997. Edge. J. Potts. (1♂). [ AMSA] ; New South Wales: EPA Survey, Murray River - Tom Groggin , MURR 24. 36 32 52S; 148 07 34E. 3-XII-1997. Edge. Eren Turak. (1♂, 2♀♀). [ AMSA]; same data: Riffle. Eren Turak. (1♂, 1♀). [ AMSA] ; New South Wales: Hawkesbury- Nepean River System, N641. 10-III-1998. Edge. J. E., C. H., & R. H. (1♂). [ AMSA] ; New South Wales: EPA Survey MRHI, Murray River - Tom Groggin , MURR 24. 36 32 52S; 148 07 34E. 24-III-1998. Edge. P. Scanes. (4♂, 3♀♀). [ AMSA] ; New South Wales: Shoalhaven River, Fossicker’s Flat. 34 49 16S; 150 11 32E. 30-IV- 1998. Edge. Natacha Waddell. (1♂). [ AMSA]; same data: 10-XII-1998. Edge. Eren Turak. (1♂). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N78. 14-I-1999. Edge. B. F. (3♂, 4♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N44. 22-I-1999. Edge. R. H. (2♂, 2♀♀). [ AMSA]; same data: Macrophytes. B. F. (2♂, 2♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N67, Wallacia Bridge , 33 51 57S; 150 38 17E. 18-V-1999. Edge. Ecology Lab. (1♂, 1♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Macquarie Grove Bridge. 34 02 36S; 150 41 36E. 19-V-1999. Edge. Ecology Lab. (1♂, 3♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Douglas Park. 34 11 42S; 150 42 38E. 27-V-1999. Macrophytes. Ecology Lab. (4♂, 4♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Yarramundi Bridge, Neapean River . 33 36 53S; 150 41 52E. 28-V-1999. Edge. Ecology Lab. (10♂, 12♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N75. 2-VI-1999. Edge. Ecology Lab. (3♂, 1♀). [ AMSA] ; New South Wales: Smith St. Ex-Bridge, Hawkesbury-Nepean River System. 33 40 10S; 150 39 44E. 3-VI-1999. Edge. Ecology Lab. (6♂, 4♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, E6133, u/s Cordeaux Dam. 34 23 21S; 150 49 28E. 22-IX-1999. Edge. Ecology Lab. (1♂, 2♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Ryan’s Crossing. 34 20 15S; 150 44 40E. 22-IX- 1999. Edge. Ecology Lab. (5♂, 7♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Maldon Bridge. 34 12 08S; 150 37 58E. 23-IX-1999. Macrophytes. Ecology Lab. (1♂, 1♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Maldon Weir. 34 12 12S; 150 37 50E. 23-IX-1999. Macrophytes. Ecology Lab. (1♂). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N931, Rockford Bridge. 34 14 53S; 150 36 23E. 24-IX-1999. Edge. Ecology Lab. (1♂, 1♀). [ AMSA]; same data. Macrophytes. Ecology Lab. (2♀♀). [ AMSA] ; New South Wales: Yarramund Bridge, Hawkesbury-Nepean River System. 33 36 53S; 150 41 52E. 24- IX-1999. Edge. Ecology Lab. (1♂). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N95, Avon Road. 34 19 47S; 150 37 42E. 27-IX-1999. Edge. Ecology Lab. (8♂, 6♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, u/s Broughton’s Pass. 34 13 49S; 150 44 34E. 27-IX-1999. Edge. Ecology Lab. (1♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Wallacia Bridge. 33 51 57S; 150 38 17E. 27-IX-1999. Edge. Ecology Lab. (3♂, 2♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, Douglas Park. 34 11 42S; 150 42 38E. 6-X-1999. Edge. Ecology Lab. (2♂, 1♀). [ AMSA]; same data. Macrophytes. Ecology Lab. (1♂, 1♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, E619, u/s Avon Dam. 34 28 04S; 150 42 52E. 6-X-1999. Edge. Ecology Lab. (3♀♀). [ AMSA] ; New South Wales: Hawkesbury-Nepean River System, N89, Douglas Park. 34 11 42S; 150 42 38E. 6-X-1999. Rock. Ecology Lab. (1♂). [ AMSA] ; Queensland: Ferny Grove, Brisbane. 27-V-1962. Creek, waterhole. A. W. Sweeney. (1♂, 3♀♀). [ USNM] ; Queensland: Brisbane, University , Queensland. 29-IX-1980. M. Pion & C. Reichart. (9♂, 13♀♀). [ USNM] ; South Australia: Musgrave Ranges, Ernabella Creek at Ernabella. 7-V-1983. W. Zeidler. (2♂, 2♀♀). [ SAMA] ; South Australia: Sturt Gorge. 25-IV-1987. Creek. C. Watts. (1♂, 2♀♀). [ SAMA] ; Tasmania: Derwent River , 2 km NW Derwent Bridge. 7-XI-1972. A. Neboiss. (3♂, 9♀♀). [ MVMA] ; Tasmania: Bronte Lagoon. 8- XI-1972. A. Neboiss. (4♂, 7♀♀). [ MVMA] ; Tasmania: Derwent River , 2 km NW Derwent Bridge. 12-XI-1972. A. Neboiss. (1♂, 1♀). [ MVMA] ; Victoria: Clunes. 9-VII-1954. A. Neboiss. (2♂). [ MVMA] ; Victoria: Gembrook Res. 9-III-1963. (3♀♀). [ MVMA] ; Victoria: Yarra Flats, Nealesville. 14-X-1965. A. Neboiss. (2♂, 4♀♀). [ MVMA] ; Victoria: Hattah Lakes. 30-VIII-1966. A. Neboiss. (1♂). [ MVMA] ; Victoria: Deadman’s Creek, Keera Hstd. Chowilla Project. 27-IV-1967. (3♂, 1♀). [ MVMA] ; Victoria: Walpolla Creek, Keera Hstd , 35 mi W of Mildura. 27-IV-1967. R. Blackwood. (3♂, 3♀♀). [ MVMA] ; Victoria: Nampoo Billabong , 90 km WNW Mildura. IX-1967. R. Blackwood. (1♂, 8♀♀). [ MVMA] ; Victoria: Berribee Punt Lindsay LS Site , 100 km W Mildura. 22- XI-1967. R. Blackwood. (10♂, 14♀♀). [ MVMA] ; Victoria: Mt. Emu Creek , 2 miles E Terang. 2-XI-1970. B. J. Smith. (3♂, 3♀♀). [ MVMA] ; Victoria: Terang . 2-XI-1970. B. J. Smith. (2♀♀). [ MVMA] ; Victoria: Tarago River , 7 km W Neerim. 1-III-1972. A. Neboiss. (1♂). [ MVMA] ; Victoria: Buffalo River, Dandongadale. 12-IV- 1972. A. Neboiss. (14♂, 12♀♀). [ MVMA] ; Victoria: Avoca River Xroads, 13 km NW Lexton. 2-X-1972. B. J. Smith & R. J. Plant. (1♂). [ MVMA] ; Victoria: Myrtleford , Eucalyptus spp. 23-I-1973. MV Trap. A. Neboiss. (1♂, 1♀). [ MVMA] ; Victoria: Dartmouth Survey, Larsen Creek, Locality N. 12-II-1973. (2♂, 3♀♀). [ MVMA] ; Victoria: Dartmouth Survey, Dart River . 2-III-1973. Melb. Uni. Bot. (2♂). [ MVMA] ; Victoria: Dartmouth Survey, Mitta Mitta River, Locality MB. 6-III-1973. (2♂, 2♀♀). [ MVMA] ; Victoria: Werribee River, Ballan. 10- VI-1976. A. Calder. (8♂, 10♀♀). [ MVMA] ; Victoria: Rocklands Res. , 3 km SSW of Glenisla. 8-III-1977. A. Calder. (6♂, 4♀♀). [ MVMA] ; Western Australia: Woodstock Stn. , WS9. 21 37 00S; 119 01 24E. 24-IX-1988. B. P. Hanich. (18♂, 24♀♀). [ WAMP] .

Figures

FIGURE 41. a–c. Micronecta batilla Hale (= M. annae Kirkaldy), holotype and dorsal habitus. a–b. Locality and type labels. c. Holotype specimen, male, dorsal habitus. d–e. M. annae, brachypterous form. d. Male. e. Female. Scale bars = 0.5 mm.

FIGURE 42. Micronecta annae Kirkaldy, brachypterous form. a. Male. b. Female. Scale bars = 0.5 mm.

FIGURE 43. Micronecta annae Kirkaldy, dorsal habitus. a–b. Brachypterous form. a. Male. b. Female. c–d. Macropterous form. c. Male. d. Female. Scale bars = 0.5 mm.

FIGURE 44. Micronecta annae Kirkaldy. a. Prothoracic lobe, female. b. Metaxyphus. c. Left male foreleg, posterior aspect. d. Left male pala (tarsus), anterior aspect, palar claw folded in. e. Prestrigilar flap of abdominal segment V. f. Sternal process of abdominal segment VII. g. Free lobe of abdominal tergum VIII. Scale bars = 0.1 mm.

FIGURE 45. Micronecta annae Kirkaldy. a. Metaxyphus. b. Left male pala, anterior aspect with palar claw everted. c. Free lobe of abdominal tergite VIII. d. Left paramere, medio-lateral aspect, right paramere, dorsal aspect. e. Left paramere, dorsal aspect. f. Right paramere, lateral aspect, left paramere, medio-ventral aspect. g. Right paramere, lateral aspect. h. Pars stridens processus, base of right paramere.