Trimorus granulatus, Komeda & Mita & Yamagishi, 2018
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publication ID |
https://dx.doi.org/10.3897/jhr.63.23671 |
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publication LSID |
lsid:zoobank.org:pub:F446E760-99FE-499D-AF5E-BF298735A6C6 |
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persistent identifier |
https://treatment.plazi.org/id/6694BCFA-36A2-4C18-A1D4-13E8381C50F7 |
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taxon LSID |
lsid:zoobank.org:act:6694BCFA-36A2-4C18-A1D4-13E8381C50F7 |
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treatment provided by |
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scientific name |
Trimorus granulatus |
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sp. n. |
Trimorus granulatus sp. n.
Figs 2 View Figure 2 ; 5 View Figure 5 ; 6 View Figure 6
Diagnosis.
Head globular. Frons granulate. Eyes small. Mandible subtridentate, with anterior and posterior large teeth and median small tooth. Mesoscutum and mesoscutellum flat, granulate. Postacetabulum granulate-smooth. In female, A2 and A3 longest among A2-6; fore and hind wings short, narrow, beyond anterior edge of metasoma; T1 without horn; T3 shallowly punctate with dense setae. In male, A5 about 2.3 times as long as wide, about 1.5 times as long as tyloid; fore and hind wings long, narrow, exceeding to apical metasoma; T3 weakly punctate-smooth.
Description.
Female (n = 3): Length = 0.90-0.95 mm (m = 0.93).
Color (Fig. 2A, B View Figure 2 ). Body dark brown; A1-2, legs, T1 light brown.
Head globular. FCI = 1.13-1.16 (m = 1.14); LCI = 1.28-1.35 (m = 1.32); DCI = 1.48-1.52 (m = 1.50); HW/IOS = 1.33-1.40 (m = 1.37); head about 1.3 times as wide as mesosoma (HW/TSL = 1.24-1.32, m = 1.29). Frons (Fig. 5A View Figure 5 ) granulate with dense setae; frontal patch absent; central keel present ventrally; antennal scrobe small, smooth, without setae; interantennal process (Fig. 2B View Figure 2 ) rounded without setae. Vertex granulate with dense setae; POL as long as OOL (POL/OOL = 0.90-1.11, m = 1.00); OOL about 1.9 times as long as LOL (OOL/LOL = 1.80-2.00, m = 1.87); hyperoccipital carina absent; vertex patch absent. Eyes small with dense setae. Malar region costate; facial striae expanding to bottom level of eye; orbital carina extending to top level of eye. Gena granulate with dense setae; genal patch absent. A1 (Fig. 5B View Figure 5 ) about 5.6 times as long as radicle (A1/r = 5.50-5.75, m = 5.58), about 22.3 times as long as A6 (A1/A6 = 22.00-23.00, m = 22.33), about as long as clava (A1/cl = 1.05-1.10, m = 1.00); A2-3 same length, 5 times as long as A6 (A2/A6 = 5.00; A3/A6 = 5.00); A4 about 3.3 times as long as A6 (A4/A6 = 3.00-4.00, m = 3.33); A5 as long as A6 (A5/A6 = 1.00). Mandible tridentate; anterior and posterior tooth same length, median tooth shorter.
Mesosoma. Pronotal suprahumeral sulcus sulcate-foveolate; epomial carina weakly present; cervical pronotal area granulate with dense setae; lateral pronotal area smooth. Mesoscutum (Fig. 5C View Figure 5 ) about 1.6 times as wide as long (TSL/ML = 1.50-1.61, m = 1.56), flat, granulate, with dense setae; mesoscutal suprahumeral sulcus weakly present; mesoscutal humeral sulcus weakly present; antero-admedian line absent; notauli weakly present, expanding to half level of mesoscutum; inter notaular area granulate with dense setae; lateral notaular area granulate with dense setae. Mesoscutellum about 2.4 times as wide as long (SW/SL = 2.25-2.50, m = 2.36), flat, granulate, with dense setae; scutoscutellar sulcus foveolate; axillular carina present; mesoscutellum without median spine; posterior scutellar sulcus foveolate. Femoral depression (Fig. 5D View Figure 5 ) smooth; mesopleural carina present; anterior rows of foveae of mesopleural carina present; posterior rows of foveae of mesopleural carina foveolate dorsally; postacetabular sulcus foveolate; postacetabulum granulate-smooth with dense setae; postacetabular patch absent; sternaulus absent; mesepimeral sulcus foveolate; speculum wide, transversely sulcate; prespecular sulcus foveolate; transpleural sulcus absent. Metanotal trough foveolate; metascutellum rugulose-foveolate; metascutellar carina unclear; metanotal spine weakly present, short, blunt. Metapleural sulcus present; dorsal metapleural areas smooth; ventral metapleural areas smooth; paracoxal sulcus foveolate; metapleural epicoxal sulcus present; metapleural epicoxal carina completely present; metapleural triangle smooth; prespiracular propodeal area narrow; lateral propodeal carina present; lateral propodeal area sulcate; metasomal depression sulcate; plica absent; posterior propodeal projection weakly present; plical area narrow, with dense setae. Legs (Fig. 2A, B View Figure 2 ) elongate. Fore wing (Fig. 2A, B View Figure 2 ) short, narrow, beyond anterior edge of metasoma. Hind wing short, narrow, beyond anterior edge of metasoma.
Metasoma. T1 about 0.6 times as long as T1+T2 length (T1W/T1+T2L = 0.50-0.59, m = 0.55), longitudinally costate. T2 longitudinally costate in anterior, shallowly punctate with dense setae in posterior; basal depressions on T2 present; lateral patch of T2 absent. T3 (Fig. 5E View Figure 5 ) about 1.2 times as wide as long (T3W/T3L = 1.10-1.22, m = 1.16), about 1.2 times as wide as mesoscutum (T3W/TSL = 1.18-1.22, m = 1.20), shallowly punctate with dense setae; basal depressions on T3 absent; lateral patch of T3 absent; posterodorsal patch of T3 absent; apical setae on T3 absent. S3 (Fig. 5F View Figure 5 ) shallowly punctate with dense setae. T4 shallowly punctate with dense setae; median patch on T4 absent; lateral patch of T4 absent. T5 shallowly punctate with dense setae; lateral patch of T5 absent. T6 shallowly punctate with dense setae; lateral patch of T6 absent.
Male (n = 3): Length = 0.93-1.00 mm (m = 0.95). FCI = 1.17-1.27 (m = 1.22); LCI = 1.32-1.36 (m = 1.34); DCI = 1.55-1.73 (m = 1.64); HW/IOS = 1.42-1.46 (m = 1.45); HW/TSL = 1.17-1.27 (m = 1.24). Central keel (Fig. 6A View Figure 6 ) present, incom plete dorsally; antennal scrobe larger than female, smooth, without setae. POL/OOL = 0.89-1.00 (m =0.96); OOL/LOL = 1.80-2.25 (m = 2.10). Eyes larger than female. Orbital carina extending to bottom level of eye. A1/r = 4.00-4.40 (m = 4.20); A5 (Fig. 6B View Figure 6 ) 2.3 times as long as wide (A5L/A5W = 2.25), about 1.5 times as long as tyloid (A5L/ty = 1.50). TSL/ML = 1.36-1.50 (m = 1.44). SW/SL = 2.00-2.20 (m = 2.13); notauli (Fig. 6C View Figure 6 ) present, expanding to 3/4 levels of mesoscutum. Mesoscutellum shallowly punctate-smooth with dense setae. Anterior rows of foveae of mesopleural carina (Fig. 6D View Figure 6 ) foveolate dorsally, smooth ventrally; postacetabulum granulate-smooth with sparse setae; speculum wide, smooth. Metapleural sulcus absent; paracoxal sulcus foveolate dorsally, absent ventrally; metapleural epicoxal sulcus absent; lateral propodeal area foveolate; metasomal depression foveolate; plica present in posterior; plical area with dense setae. Legs (Fig. 2C, D View Figure 2 ) elongate. Fore wing (Fig. 2C, D View Figure 2 ) long, narrow, exceeding to apical metasoma, as wide as mesoscutum (TSL/WW = 0.94-1.04, m = 1.00); marginal vain about 3.5 times as long as stigmal vein (m/st = 3.33-3.67, m = 3.53). Hind wing long, narrow, exceeding to apical metasoma, 1.5 times as wide as length of marginal cilia at widest point (HWW/HWS = 1.40-1.50, m = 1.47). T1W/T1+T2L = 0.60-0.63 (m = 0.61). T3W/T3L = 1.11-1.23 (m = 1.18); T3 (Fig. 6E View Figure 6 ) narrower (T3W/TSL = 1.03-1.07, m = 1.04), weakly punctate-smooth with sparse setae; posterodorsal patch of T3 present, rugulose. S3 (Fig. 6F View Figure 6 ) smooth with dense setae; S3 setae sparser than female. T4 smooth with dense setae; T4 setae sparser than female.
Material examined.
Holotype: Fukuoka pref.: Fukuoka city, Mt. Tachibana, 2. X. 1993, H. Honda leg., 1♀ (YPT) [ELKU] . Paratypes: Same locality as the holotype , 2. X. 1993, H. Honda leg., 5♂ (YPT) [ELKU]; 16. X. 1993, 7♂ 2♀ (YPT) [ELKU]; 23. X. 1993, 5♂ 5♀ (YPT) [ELKU]; 30. X. 1993, 6♂ 1♀ (YPT) [ELKU]; 6. XI. 1993, 7♂ 2♀ (YPT) [ELKU]; 14. XI. 1993, 8♂ 6♀ (YPT) [ELKU]; 20. XI. 1993, 6♂ 6♀ (YPT) [ELKU]; 27. XI. 1993, 3♂ 2♀ (YPT) [ELKU].
Distribution.
Japan (Kyushu: Fukuoka).
Etymology.
The species name refers to the sculpture on frons and mesoscutum.
Remarks.
Among eastern Palearctic Trimorus species, the female of this species is similar to T. amesis Kozlov & Kononova, 2001 and T. mirandus Kozlov & Kononova, 2001 in shortened wings at most reaching posterior margin of mesosoma. But it differs from T. amesis in small body size ( T. granulatus about 0.9 mm; T. amesis about 1.4 mm) and sculpture of T3 (shallowly punctate in T. granulatus , Trimorus coriaceus in T. amesis ), and from T. mirandus in sculpture of frons, mesoscutellum and T3 (frons and mesoscutellum are granulate (Fig. 5A, C View Figure 5 ) and T3 is shallowly punctate (Fig. 5E View Figure 5 ) in T. granulatus ; all of them are smooth in T. mirandus ). In addition, this species differs from T. coriaceus sp. n. in the shape of head (globular in T. granulatus (Fig. 2A, B View Figure 2 ), flat in T. coriaceus (Fig. 1A, B View Figure 1 )) and sculpture of frons (granulate in T. granulatus (Fig. 5A View Figure 5 ); coriaceous in T. coriaceus (Fig. 4A View Figure 4 )) and T3 (shallowly punctate in T. granulatus (Fig. 5E View Figure 5 ); densely punctate-imbricate in T. coriaceus (Fig. 4E View Figure 4 )). Male of T. granulatus is similar to T. bisulcatus Kieffer, 1908 in ratio of length and width of antenomeres. The new species differs from T. bisulcatus in body size ( T. granulatus is about 1 mm; T. bisulcatus is about 1.7-2 mm) and body sculpture.
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