Nannaria asta, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

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scientific name

Nannaria asta

sp. nov.

Nannaria asta   sp. nov. Figs 90 View Figure 90 , 91 Vernacular name: "The Crawfish Valley Twisted-Claw Millipede" View Figure 91

Material examined.

Holotype: United States - Virginia • ♂; Wythe County, Crawfish Valley, Channel Rock , ca. 1.5 mi down trail from Strawberry Rd. end; 36.9585°N, - 81.3189°W; elev. 770 m; 24 Mar. 2017; hand collected; C. Harden leg.; VTEC MPE02419 View Materials . GoogleMaps  

Paratypes: United States - Virginia • 3 ♂♂; Wythe Co. Crawfish Valley ; 36.9811°N, - 81.2934°W; elev. 715 m; 24 Mar. 2017; C. Harden leg.; VTEC MPE02421 View Materials , VMNH, MPE02429 View Materials , 2430. For detailed collection data see Suppl. material 7 GoogleMaps   .


Adult males of Nannaria asta   sp. nov. are distinct from other Nannaria   and the sympatric N. solenas   sp. nov. and N. aenigma   based on the following combination of characters: Gonopods. Gonopodal acropodite gently curving medially, not straight as in N. solenas   sp. nov. Distal zone curving dorsomedially, tip rounded, directed caudally with small lateral flange (Fig. 90A View Figure 90 ), not rectangular, directed medially at 90° angle with acropodite and slight cephalically-directed upturn at terminal edge as in N. solenas   sp. nov., and not sinuous, without flange as in N. aenigma   . Acropodite with slight swelling on inner margin (Fig. 90B View Figure 90 , red triangle) and dimple on outer margin (Fig. 90A View Figure 90 , red arrow). Prefemur with stout, acicular prefemoral process, arising dorsomedially from projected, cephalically-curving prefemoral spine, not arising from prefemur as in N. solenas   sp. nov. Telopodite basal zone with small lateral bulge, lacking in N. solenas   sp. nov. Color. Tergites with orange paranotal spots (Fig. 91 View Figure 91 ). Black background. Dorsum of collum smooth with orange margin.


♂ holotype (VTEC, MPE02419): BL = 26.2, CW = 4.1, IW = 2.1, ISW = 0.9, B11W = 4.9, B11H = 3.1.


No known variation.


Known only from southwestern Virginia (Wythe and Bland counties; Suppl. material 7; Fig. 126 View Figure 126 ). The distribution of N. asta   sp. nov. is restricted to the Crawfish Valley area and south of Wytheville, Virginia. Distribution area: 171 km2; status: MRE.


The majority of Nannaria asta   sp. nov. individuals were found at night, walking along the side of a path on top of predominantly pine litter, a notably odd behavior for Nannaria   , which typically remain beneath the leaf litter and are more cryptic in their behavior.


The specific epithet is an arbitrary combination of letters derived from the Infraorder Astacidea  (which includes crayfish). It is to be treated as a noun in apposition.

Type locality.

United States, Virginia, Wythe County, Crawfish Valley, Channel Rock, ca. 1.5 mi down trail from Strawberry Rd. end, 36.9585°N, - 81.3189°W.