Rhaucus florezi, García & Kury, 2017

García, Andrés F. & Kury, Adriano B., 2017, Taxonomic revision of the Andean harvestman genus Rhaucus Simon, 1879 (Arachnida, Opiliones, Cosmetidae), Zootaxa 4338 (3), pp. 401-440: 426-430

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Rhaucus florezi

new species

Rhaucus florezi   new species

( Figs 1e View FIGURE 1 , 15–17 View FIGURE 15 View FIGURE 16 View FIGURE 17 , 18e View FIGURE 18 , 23 View FIGURE 23 )

Etymology. The species name is a genitive noun in honor to the Colombian arachnologist Eduardo Flórez, for his contributions to the study of Colombian arachnids.

Type material. Holotype ♂ (ICN-AO-1187), COLOMBIA, Boyacá Department: Tipacoque, Vereda La Calera [06°23'N 72°43'W], 2800 m, 1–5.iv.2013, M. Medrano leg. GoogleMaps   – Paratype: 1♂ (ICN-AO-1187.1), same data as the holotype GoogleMaps   .

Distribution and habitat. COLOMBIA, Boyacá Department: Pisba, Susacón, Tipacoque, Villa de Leyva. Between 2800 and 3500 m a.s.l., in Northern Andean páramo and Magdalena Valley montane forests ecoregions ( Figs 23 View FIGURE 23 , 24a View FIGURE 24 ).

Diagnosis. Leg IV coarsely granulated, with one prolateral and one retrolateral row of robust tubercles ( Fig. 16e View FIGURE 16 ; contrasting with the weakly granulated leg IV of R. quinquelineatus   ); a pair of short mammilliform spines in area III ( Fig. 15c View FIGURE 15 ; contrasting with the high domed spines of R. robustus   , R. serripes   and R. quinquelineatus   , and differing from R. vulneratus   by having sharper tips).

Description. Male holotype (ICN-AO-1187)

Measurements: Holotype: CL=2.1, CW=4.1, AL=2.8, AW=5.2, BaCh=0.9, FeLIII=4.1, FeL IV=5, TiL III= 2.8, TiL IV= 4. Intraespecific variation of body and appendages measurements in Table 1.

Dorsum ( Figs 1e View FIGURE 1 , 15a, 15c View FIGURE 15 , 16a –b View FIGURE 16 , 18e View FIGURE 18 ): Dorsal scutum alpha   type, slightly granulate ( Fig. 15a View FIGURE 15 ), with five yellow transverse curved lines that vary in length and shape and a reticulate pattern each side of ocularium ( Figs 15a View FIGURE 15 , 18e View FIGURE 18 ). Abdominal scutum widest at level of groove II; areas I –IV with some granules. Carapace with some granules on the anterior region ( Fig. 15a View FIGURE 15 ). Ocularium low without median depression, with some granules ( Figs 15c –e View FIGURE 15 , 16a View FIGURE 16 ). Lateral margins of DS with a few granules on the medial region ( Figs 16a –b View FIGURE 16 ). Mesotergum delimited, divided into four areas: area I with a pair of short paramedian tubercles; area II with a pair of tiny tubercles, laterally displaced; area III with a pair of paramedian, mammilliform, granulated tubercles; area IV with a pair of small paramedian tubercles ( Figs 15c View FIGURE 15 , 16a –b View FIGURE 16 ). Posterior border of scutum sub-straight and with a row of tubercles. Free tergites I –III with a row of tubercles ( Figs 16a –b View FIGURE 16 ).

Venter ( Fig. 15b View FIGURE 15 ): Stigmatic area with a few granules. Stigmata large, oval and transverse. Coxa I with two parallel rows of tubercles, increasing in size distally, and with an anterior smooth space for the pedipalp to rest; coxa II slightly longer than coxa I, granulate; coxa III longer than I and II, granulate, with posterior margin sigmoid; coxa IV strongly backward, granulate. Genital operculum slightly granulate.

Pedipalps ( Figs 17a –c View FIGURE 17 ): Coxa dorsally and ventrally with one meso-distal granule. Trochanter with one dorsal short tubercle and one ventral tubercle; femur compressed, dorsally with a row of aligned triangular and acuminate setiferous tubercles of different sizes –not reaching the distal portion- and ventrally with a row of rounded tubercles (the two basal-most tubercles fused, the three distal-most tubercles smaller than the others). Patella distally depressed with dorsal granules, the distal taller; tibia depressed, spoon-shaped, dorsally granulate, with lateral borders tuberculate and some spiniferous tubercles at distal portion; inner surface slightly striated ( Figs 17a – b View FIGURE 17 ).Tarsus long, conical, with some dorsal granules and ventrally with two rows of thickened setae ( Fig. 17c View FIGURE 17 ).

Chelicera ( Figs 15 View FIGURE 15 , 16a –b View FIGURE 16 , 17d –e View FIGURE 17 ): Chelicera swollen. Basichelicerite rectangular, with a few granules; the proximal border with some tubercles, the dorso-ectal larger; ectal face with two basal small tubercles, one mediumsized tubercle slightly curved and a group of distal tubercles (one of them larger and curved); mesal face with some granules and one large distal tubercle ( Fig. 17d View FIGURE 17 ). Hand with one tubercle near the joint of the movable finger. Fixed finger with the inner surface finely grooved. Movable finger with one conical, sub-basal tooth and with the distal portion of the inner surface dentate ( Fig. 17e View FIGURE 17 ).

Legs ( Figs 15a –b View FIGURE 15 , 16d –e View FIGURE 16 ): Coxae I –II dorsally with one posterior process; coxa III with one proximal retrolateral tubercle; coxa IV coarsely granulate ( Figs 15a View FIGURE 15 , 16a View FIGURE 16 ), with a prominent anterolateral clavi inguines and a posterolateral multicapitate apophysis ( Fig. 16a View FIGURE 16 ). Trochanters I –II dorsally with some minute granules and ventrally with some tubercles; trochanter III with some dorsal and ventral tubercles, and with one medium-sized, proximal prolateral tubercle and one retrolateral tubercle; trochanter IV with some dorsal and ventral tubercles, and with one large, proximal prolateral tubercle and one medium-sized retrolateral tubercle ( Figs 16d –e View FIGURE 16 ). Femora I –IV sigmoid, granulated; III –IV ventrally with two longitudinal rows of tubercles along the entire length, and one prolateral and one retrolateral row of large tubercles (the retrolateral row of leg IV has the tubercles closer together compared to leg III) ( Figs 16d –e View FIGURE 16 ). Patellae I –IV dorsally granulate, curved (patella IV with large tubercles at distal border) ( Fig. 16e View FIGURE 16 ). Tibiae I –IV granulate; tibiae III –IV sub-straight, with two longitudinal rows of tubercles along its length (in tibia III the prolateral row has smaller tubercles than the retrolateral row); tibia IV with tubercles of similar size (the distal tubercle of retrolateral row is fused) ( Figs 16d –e View FIGURE 16 ). Metatarsus I –IV with one ventrodistal setae. Tarsi I –II with one smooth claw; tarsi III –IV with 2 subparallel smooth claws and tarsal process ( Fig. 16c View FIGURE 16 ). Tarsal counts: 6(3)–6(3)/13(3)–14(3)/9–9/9–9. Intraspecific variations in Table 2.

Color (in ethanol): Carapace Deep Brown (56), reticulated in the ocularium. Transverse lines in carapace Brilliant Yellow (83). Pedipalps, chelicera and coxae I –IV Strong Brown (55), reticulated. Trochanters I –IV Strong Yellowish Brown (74). Legs I –IV Strong Reddish Brown (40).

Male genitalia ( Figs 17f –h View FIGURE 17 ): VP of penis subrectangular with slightly concave distal border; VP with two lateral, elongated, dense patches of type 4 microsetae, separated by a longitudinal space ( Fig. 17h View FIGURE 17 ). VP with two apical MS C, curved and laterally inserted; two MS D, the most distal large and straight, following the same lateral row as MS C, and the other smaller, inserted dorso-laterally on the middle third of VP; two MS A straight and laterally inserted, near D2 ( Figs 17f –g View FIGURE 17 ); two small MS E volcano-shaped on the ventral face; one small MS B volcano-shaped inserted laterally at the base of the VP ( Fig. 17f View FIGURE 17 ). Glans mostly smooth; stylus long, wattle barbels long ( Fig. 17g View FIGURE 17 ), finger-like process rounded, enlarged ( Fig. 17f View FIGURE 17 ).

Female. Similar to male, but differs by having anterior part of carapace narrower and coda divergent; tubercles of area I and III shorter than male; legs III –IV thinner; tubercles in ventral rows of legs III –IV smaller; chelicerae not hyperthelic ( Figs 15d –e View FIGURE 15 ).

Other material examined. 12 specimens (7 males, 5 females) in 11 lots. COLOMBIA   . Boyacá Department: Villa de Leyva, 1♂ (ICN-AO-294), S.F.F. Iguaque sector Carrizal [05°41'N 73°27'W], 2800 m, 23.i.1998, L. Ballesteros leg. GoogleMaps   ; 1♂ (ICN-AO-297), S.F.F. Iguaque sector Carrizal [05°41'N 73°27'W], 2900 m, 9.vi.2001, E. Flórez leg. GoogleMaps   ; 1♂ 1♀ ( MNRJ 8884 View Materials ), S.F.F. Iguaque sector Carrizal [05°41'N 73°27'W], 2800 m, 31.x.2004, E. Flórez, G. Giribet & M. Porras leg. GoogleMaps   ; 1♀ (IAvH-3000180/ IAvH-I-6), S.F.F. Iguaque, Cabaña Chaina [05°25'N 73°27'W], 9.vi.2001 GoogleMaps   ; 1♀ (IAvH-I-10), S.F.F. Iguaque , lagoon shore [05°38'N 73°29'W], 3450 m, 13.ix.1998, E. González leg. GoogleMaps   ; 1♀ (IAvH-3000007/IAvH-I-13), S.F.F. Iguaque , lagoon shore [05°38'N 73°31'W], 3340 m, 14.x.1998, E. González leg. GoogleMaps   ; Pisba, 1♂ (ICN-AO-1191), Páramo de Pisba [5°44'N 72°29'W], 24.xi.1974 GoogleMaps   ; Susacón, 1♀ (IAvH-3000137/ IAvH-I-5), Susacón river, 20 km from Susacón [06°10'N 72°46'W], 3 135 m, 23.ii.1999, E. González leg. GoogleMaps   ; Aquitania, 1♂ (IAvH-3000136/IAvH-I-9), Los Curies Páramo ( Guarne ) [05°25'N 72°56'W], 3500 m, 8.ii.1999, E. González leg. GoogleMaps