Phylloicus maculatus (Banks)

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 66-68

publication ID 10.11646/zootaxa.275.1.1

publication LSID


persistent identifier

treatment provided by


scientific name

Phylloicus maculatus (Banks)


Phylloicus maculatus (Banks) View in CoL

Figs. 64 View FIGURE 64 , 65 View FIGURE 65

Phylloicus maculatus ( Banks, 1901:369) View in CoL [Type locality: Mexico, Veracruz, Presidio; MCZ; female; in Heteroplectron View in CoL ]. — Flint 1967:19 [female, to Phylloicus View in CoL ]. — Holzenthal 1988:72 [distribution].

Phylloicus maculatus View in CoL is distinguished by its large size, wing pattern, and by having short, single­lobed lateral coremata, without modifications of tergite IV ( Fig. 64F View FIGURE 64 ). The wing pattern of this species is identical to that of P. elegans View in CoL and lituratus View in CoL . The female genitalia are indistinguishable from those of the latter two species.

Adult. Forewing length 13.415. 4 mm, n = 31.

Head dark brown, setal warts very dark. Maxillary palps dark brown, covered with golden brown setae on basal three segments. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown, mesoscutellum golden brown; ventrolateral thorax golden. Femora golden; tibiae golden; tarsi dark brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, ivory. Hind wing basal brush present in male and female; female brush short.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with expanded lateral flanges, lateral coremata; lateral coremata single­lobed and short, cylindrical. Tergum V without sclerotized modifications ( Fig. 64F View FIGURE 64 ). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched ( Fig. 64A, C View FIGURE 64 ). Tergum IX with paired mesal ridges extending 1/2 length of segment; posterior margin with round narrow mesal projection ( Fig. 64B View FIGURE 64 ); lateral ridge absent; dorsal pleural setae approximately 5, ventral pleural setae approximately 8 ( Fig. 64A View FIGURE 64 ). Preanal appendage longer than tergum X, but less than 11/2 times length, widest apically, setae filamentous, longer than appendage ( Fig. 64A, B View FIGURE 64 ). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes long, length at least twice diameter; apex, in lateral view, rounded, in dorsal view, entire ( Fig. 64A, B View FIGURE 64 ). Harpago rounded; peglike setae many, apical ( Fig. 64A, C View FIGURE 64 ). Phallic endotheca with paired basolateral lobes, basolateral lobes tapered apically; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 64D, E View FIGURE 64 ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII ( Fig. 65C View FIGURE 65 ). Tergum IX without mesal ridge ( Fig. 65B View FIGURE 65 ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 65A View FIGURE 65 ). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes length approximately equal diameter ( Fig. 65B View FIGURE 65 ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 65A View FIGURE 65 ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized ovoid ( Fig. 65A View FIGURE 65 ).

Material examined. GUATEMALA: Izabal: Cayuga , 15°32'00”N, 88°42'00"W, 1.ii.1915, Schaus — 1 female ( NMNH) GoogleMaps ; MEXICO: Veracruz: Presidio , 19°05'00”N, 96°58'00"W, holotype female ( MCZ) GoogleMaps ; Municipio Tilapan, Arroyo Quetzalan. Los Manantiales km 5 Rta. Orizaba­Zongolica , 18°47'28”N, 97°06'05"W, 1160 m, 17.iii.2000, Bueno, Barba, & Rojas — 1 female ( UMSP) GoogleMaps ; 13.vii.2001, Bueno & Barba — 12 males, 10 females ( IBUNAM) ; — 3 males, 3 females ( UMSP) .

Distribution. Guatemala, Mexico


Smithsonian Institution, National Museum of Natural History


Museum of Comparative Zoology


University of Minnesota Insect Collection


Instituto de BiIología, Universidad Nacional Autónoma de México














Phylloicus maculatus (Banks)


Phylloicus maculatus ( Banks, 1901:369 )

Holzenthal, R. W. 1988: 72
Flint, O. S., Jr. 1967: 19
Banks, N. 1901: 369
GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF