Phylloicus perija, Prather, 2003

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 86-87

publication ID 10.11646/zootaxa.275.1.1

publication LSID


persistent identifier

treatment provided by


scientific name

Phylloicus perija

sp. nov.

Phylloicus perija , new species

Figs. 88 View FIGURE 88 , 89 View FIGURE 89

Phylloicus perija is most similar to P. abdominalis and P. pirapo . However, it differs in the contorted shape of the tergum IV lateral sclerite, and the setose posterior process of tergum V ( Fig. 88F View FIGURE 88 ) and in the forewing pattern. The forewing lacks the white spot on the nygma, and the wing bands are golden rather than white or ivory.

Adult. Forewing length 10.1­12.8 mm, n = 24.

Head chestnut brown. Maxillary palps chestnut brown. Antenna twice forewing length; chestnut brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax golden brown; ventrolateral thorax golden brown. Legs golden brown. Tibial spur formula 2,4,4. Forewing flat; chestnut brown; with two transverse bands; proximal band golden, reaching posterior wing margin, at least 1/2 width of wing; distal band golden, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, golden; small patch of golden setae at proximal end of wing coupling hairs. Hind wing basal brush present in male and female.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum III with anterolateral flanges. Tergum IV with paired posterior processes and paired lateral sclerites, mesal coremata and lateral coremata; posterior process truncate; lateral sclerite contorted, setose apically; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata bilobed, without setae, capable of inflation greater than length of segment V. Tergum V emarginate posteriorly, with setose posterior processes ( Fig. 88F View FIGURE 88 ). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched ( Fig. 88A, C View FIGURE 88 ). Tergum IX without mesal ridge; posterior margin obtuse; thinly sclerotized anteromesally, muriculate ( Fig. 88B View FIGURE 88 ); lateral ridge absent; dorsal pleural setae approximately 8, ventral pleural setae absent ( Fig. 88B View FIGURE 88 ). Preanal appendage longer than tergum X, but less than 11/ 2 times length, widest apically, setae filamentous, longer than appendage ( Fig. 88A, B View FIGURE 88 ). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes absent; apex, in lateral view, rounded, in dorsal view, entire or notched, notch shallow ( Fig. 88A, B View FIGURE 88 ). Harpago rounded; peglike setae few, apical ( Fig. 88A, C View FIGURE 88 ). Phallic endotheca with paired basolateral lobes, basolateral lobes large and round; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 88D, E View FIGURE 88 ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII posterior margin entire; sternum VIII ( Fig. 89C View FIGURE 89 ). Tergum IX without mesal ridge ( Fig. 89B View FIGURE 89 ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 89A View FIGURE 89 ). Tergum X appendage shorter than mesal lobe, base indistinct, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes length approximately equal diameter ( Fig. 89B View FIGURE 89 ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 89A View FIGURE 89 ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite ovoid ( Fig. 89A View FIGURE 89 ).

Holotype male: VENEZUELA: Zulia: Parque Nacional Perijá, Río Negro in Toromo , 10°03'04”N, 72°42'43"W, 360 m, 15.i.1994, Holzenthal, Cressa, & Rincón ( UMSP). GoogleMaps

Paratypes: VENEZUELA: Zulia: Perijá El Tucuco, Mission El Tucuco, Río del Pelaya , 2­1/ 2 km from church, 26.iv.1981, Townes — 1 male ( NMNH) ; Parque Nacional Perijá, Río Negro in Toromo , 10°03'04”N, 72°42'43"W, 360 m, 15.i.1994, Holzenthal, Cressa, & Rincón — 5 females ( IZAM) GoogleMaps ; — 1 female, 1 male ( NMNH) GoogleMaps ; — 11 females, 4 males ( UMSP) .

Distribution. Venezuela.

Etymology. Named for the type locality, Perijá National Park, Venezuela.


University of Minnesota Insect Collection


Smithsonian Institution, National Museum of Natural History

GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF