Phylloicus bertioga, Prather, 2003

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 31-32

publication ID 10.11646/zootaxa.275.1.1

publication LSID


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scientific name

Phylloicus bertioga

sp. nov.

Phylloicus bertioga , new species

Figs. 23 View FIGURE 23 , 24 View FIGURE 24

This species can be recognized by the single ivory band on the forewing and the short preanal appendages of the male. Neither of the male specimens had the phallic endotheca fully everted, so I am unable to describe the membranous lobes.

Adult. Forewing length 7.7­8.8 mm, n = 3.

Head dark brown, with dorsomesal crest of black setae. Maxillary palps chestnut brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown, lateral margins black; ventrolateral thorax golden brown. Femora golden brown; foretibiae dark brown; mesotibiae dark brown; metatibiae dark brown; tarsi dark brown. Metathoracic leg of male without posterior fringe. Tibial spur formula 2,4,4. Forewing flat; dark brown; with single transverse band; distal band ivory, not reaching either wing margin, 1/2 width of wing or less. Hind wing basal brush absent.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII without anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with short mesal ridge extending from anterior notch; posterior margin not distinct from base of tergum X ( Fig. 23B View FIGURE 23 ); lateral ridge absent; dorsal pleural setae approximately 3, ventral pleural setae absent ( Fig. 23A View FIGURE 23 ); sternum IX with paired mesolateral ridges; sternum IX ( Fig. 23C View FIGURE 23 ). Preanal appendage less than 2/3 length of tergum X, as long as wide, rounded apically, setae long, but not filamentous or longer than appendage ( Fig. 23A, B View FIGURE 23 ). Tergum X without basal lobes; basodorsal process absent; basolateral processes short, length less than or equal to diameter; apex, in lateral view, truncate, in dorsal view, notched, notch round ( Fig. 23A, B View FIGURE 23 ). Harpago long, curving mesally and tapering apically; peglike setae tiny, mesal ( Fig. 23A, C View FIGURE 23 ). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 23D, E View FIGURE 23 ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII without anteromesal process. Tergum VIII with sparser posterolateral brush of setae ( Fig. 24B View FIGURE 24 ); sternum VIII cleft posteromesally to anterior ridge; sternum VIII ( Fig. 24C View FIGURE 24 ). Tergum IX with mesal ridge extending 1/3 length of segment ( Fig. 24B View FIGURE 24 ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with irregular, semimembranous pockets lateral to vaginal opening ( Fig. 24A View FIGURE 24 ). Tergum X appendage length equal to mesal lobe, base marked by faint suture line, apex rounded; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 24B View FIGURE 24 ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 24A View FIGURE 24 ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite ovoid; posterior end of spermatheca a tiny sclerotized sphere ( Fig. 24A View FIGURE 24 ).

Holotype male: BRAZIL: São Paulo: Bertioga , 23°51'00”S, 46°09'00"W, 5 m, 7­ 9.x.1996, Becker ( MZUSP). GoogleMaps

Paratypes: BRAZIL: São Paulo: Bertioga , 23°51'00”S, 46°09'00"W, 5 m, 7­ 9.x.1996, Becker — 1 female, 1 male ( NMNH) GoogleMaps .

Distribution. Brazil.

Etymology. Named for Bertioga, the type locality in São Paulo state.


Museu de Zoologia da Universidade de Sao Paulo


Smithsonian Institution, National Museum of Natural History

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