Dorynota (Dorynota) pugnax ( Boheman, 1854 )

Simões, Marianna V. P. & Sekerka, Lukáš, 2015, Review of the Neotropical Leaf Beetle SubgenusDorynota s. str. Chevrolat (Coleoptera: Chrysomelidae: Cassidinae: Dorynotini), The Coleopterists Bulletin 69 (2), pp. 231-254 : 246

Dorynota (Dorynota) pugnax ( Boheman, 1854) , restored status

( Figs. 32–33 , 46 )

Batonota pugnax Boheman, 1854: 161 (type locality: ‘Columbia’).

Type Material Examined. Holotype, pinned: ‘E. Coll. ∣ Chevt. [w, p, cb] ∥ Type [w, p, s circle label with red frame] ∥ 44 [g, p, s] ∥ Batonota ∣ pugionata ∣ Bhn ∣ Columbia [w, s hw by Chevrolat] ∥ 67⋅56 [w, p, sl]’ ( BMNH).

Additional Material Examined (4). COLOMBIA: ‘ Columbia’ , (1 specimen, LSC) ; PANAMA: Chiriquí: ‘ Chiriqui’ (1 specimen, DBET; 1 specimen, MMUE) ; VENEZUELA: ‘ Venezuela’ , (1 specimen, BMNH ex coll. J. S. Baly and published by Boheman (1862) as D. gladiator ) .

Diagnosis. See diagnosis under D. monoceros and Table 2.

Remarks. Boheman (1854) did not state the precise number of specimens. However, he mentioned ‘Mus. Dom. Chevrolat’ as the depository and gave single length and width measurements. In such cases, Boheman always had a single specimen, and because there is only a single specimen in the Chevrolat collection, we consider it the holotype.

The species was synonymized with D. nodosa by Spaeth (1923) based on the primary description. However, examination of the type revealed that this species is actually morphologically close to D. monoceros and not to D. nodosa . The latter distinctly differs in that it has a very short dorsal spine, which is barely longer than the width of its base, and the elytra are strongly emarginate and more protruding behind the humeral angles. Therefore, we restore D. pugnax to species status. It can be separated from D. monoceros by its shorter elytral spine, less dense and finer punctation on the elytra, and the elytra with mostly distinct intervals, while D. monoceros has very coarse and dense elytral punctation with very narrow intervals and punctures nearly touching each other.

The species is also probably found in Venezuela as the specimen reported by Boheman (1862) under D. gladiator and cited here most likely belongs to D. pugnax .

Distribution. Colombia ( Boheman 1854), Venezuela [?] ( Boheman 1862). New country record for Panama ( Fig. 46 ).

Figures

Figs. 32–43. Dorynota (s. str.) species. 32) D. pugnax, holotype, dorsal view; 33) D. pugnax, lateral view; 34) D. rileyi, paratype, dorsal view; 35) D. rileyi, lateral view and labels; 36) D. rufomarginata, holotype, dorsal view; 37) D. rufomarginata, frontal view; 38) D. rufomarginata, lateral view; 39) D. wappesi, holotype, dorsal view; 40) D. wappesi, lateral view; 41) D. yucatana, syntype, dorsal view; 42) D. yucatana, frontal view; 43) D. yucatana, lateral view.

Fig. 46. Geographic distribution of seven Dorynota (s. str.) species.