Perspicuus csincsii, Szabó & Sendi & Ősi, 2024

Szabó, Márton, Sendi, Hemen & Ősi, Attila, 2024, A new species of the vitismin cockroach genus Perspicuus Koubová, 2020 from the Late Cretaceous (Santonian) of Hungary, Zootaxa 5437 (1), pp. 105-114 : 107-109

Perspicuus csincsii sp. n.

urn:lsid:zoobank.org:act:4DFEFCC7-09C0-4FCC-B129-76DBC2F4D2AE

Fig. 2

Type material: NHMUS PAL 2023.43 .1. (holotype, male) .

Etymology: The specific name honors Szabolcs Csincsi, enthusiastic supporter of the Hungarian Dinosaur Expedition project, also good friend of author MSZ.

Type locality: Ajka-Csingervölgy [Ajka-Csinger valley], approximately 4 km SE of the city Ajka, Bakony Mountains, Hungary .

Horizon and age: Ajka Coal Formation, unknown shaft of the Ajka-Csingervölgy coal minery; Upper Cretaceous, Santonian, 86.3–83.6 Ma.

Preservation: The inclusion is incomplete. Head, thorax, left cercus and most of legs are missing. The piece of amber was embedded in epoxy resin (2010), cut and polished to make the inclusion itself more visible. Some parts of the specimen, including parts of an antennae and a maxillary palp, were destroyed. However, these body parts were photographed by Zsófia Hajdu without scale bars.

Description: Very small species, with an estimated total length of 4.75 mm. Maxillary palps with heavily setated (short-type seta) cup-like terminal palpomere ( Fig 2a,b ). Antenna filiform, antennomeres densely covered with sensillae of different lengths ( Fig. 2c,d ). Forewings (ca. 2.4 and 2.1 mm long as preserved) dark black with small pale dot/ stripe, and without pubescence ( Fig. 2e ). Abdomen (3.1 mm TL) is slightly curved dorsally with thickness extending to the central axis. Maximum thickness of abdomen is 1.3 mm. Tergites and sternites are barely recognizable. Diversification between fore, middle and hind legs in trend of protraction (from fore to hind legs), all legs are armed with long spines and spurs ( Fig. 2f,h,i ). Terminal femoral spur also present. Claws (observable only on a suggested fore leg) asymmetrical, 0.1 and 0.14 mm long ( Fig. 2g ). Arolium trapezoidal and huge 0.16 x 0.11 mm in diameter) with rounded edges ( Fig. 2g ). Cerci multisegmented, arched, very long ca. 1.45 mm as preserved) and sparsely covered by long sensillae (best preserved longest medial sensillae 0.21–0.32 mm), cercomeres flattened and trapezoid ( Fig. 2j,k ). Number of cercomeres is indistinct. Styli not visible.

Differential diagnosis: The new species can be placed within Perspicuus on the basis of cup-like terminal palpomere of the maxillary palps (all Umenocoleidae ), extremely long cerci (synapomorphy with Alienopterix and Jantaropterix ), fully carinated legs (synapomorphy with Vitisma ) and black forewing with pale dots/stripes (more elongate compared with Vitisma ). The new species differs clearly from P. pilosus and P. vrsanskyi by its size, its more extensive dark colouration and the absence of pubescence on the wings.

Autapomorphies: Possible lack of pubescence (as Jantaropterix plesiomorphically has pubescence).

Figures

FIGURE 2. Perspicuus csincsii sp. n. a) photo of maxillary palp. b) line drawing of maxillary palp. c) photograph of partially preserved antenna. d) line drawing of antennal segments. e) habitus in dorsal view. f) indicated fore leg under the ventral side of the body. g) line drawing of the claws and arolium of the same leg. h) part of hind leg in dorsal aspect. i) line drawing of suggested middle leg in ventral aspect. j) cercus in dorsal aspect. k) line drawing of the cercus. Figures (a) and (b) have no scale bars, as these parts were destroyed during preparation and are no longer available for further examination.