Pollicaria mouhoti monochroma Kongim & Panha, 2013

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pollicaria mouhoti monochroma Kongim & Panha, 2013

Figs 6A, B , 8C

Pollicaria myersii [non Haines]- Solem 1966: 13, on limestone outcrops 20 km. east of Wang Sapung [District] near Loei [Province], Thailand.

Pollicaria mouhoti monochroma Kongim & Panha in Kongim et al. 2013: 32, 33, figs 2c, 4j, k, 6c. Type locality: limestone outcrop with dry forest at Wat Tam Pha Bing, Wungsapoong District, Loei Province, Thailand. BEDO 2017: 86. Sutcharit et al. 2018: 156.

Type material examined.

Holotype CUMZ 1577 and paratypes CUMZ 1548 (9 shells) figured in Kongim et al. (2013: figs 4j, k). Paratypes CUMZ 1562 (85 shells and 10 specimens in ethanol; Figs 6B , 8C ) from Tam Pha Bing Temple, Wungsapoong District, Loei Province, 11 June 2013.

Other material examined.

CUMZ 12182 (3 juvenile shells; Fig. 6A ) from Tham Suea Lueang Temple , Mueang Loei District, Loei Province, 1 Sept. 2020 . CUMZ 12183 (4 shells) from Tham Pha Poo, Mueang Loei District , Loei Province, 1 Sept. 2020 . CUMZ 12184 (3 adult shells and 2 juvenile shells) from Phu Pha Lom , Mueang Loei District, Loei Province, 1 Sept. 2020 . CUMZ 12185 (3 adult shells and 7 juvenile shells) from Tham Pha Phung Temple , Wang Saphung District, Loei Province, 2 Sept. 2020 . CUMZ 12186 (3 adult shells and 3 juvenile shells) from Pa Phaya Temple, Suwannakhuha District , Nong Bua Lam Phu Province, 31 Aug. 2020 .

Diagnosis.

Shell height <35 mm. Shell entirely black to purple. Dorsal side of last whorl with bold wrinkles. Aperture almost round, shallow posterior angled groove present; apertural lip expanded, yellow to pale orange. Umbilicus narrow.

Differential diagnosis.

This subspecies is different from the nominotypical subspecies by a smaller shell size, an entirely black to purple shell, and a distinct karyotype pattern of (7m+3sm+2st+1t) ( Kongim et al. 2009, 2013).

Distribution.

Loei and Nong Bua Lam Phu provinces, northeastern Thailand ( Kongim et al. 2013).

Remarks.

DNA data are required to demonstrate whether P. m. monochroma is distinct from the nominotypical subspecies and should be elevated to specific status.

Figures

Figure 6. A, B Pollicaria mouhoti monochroma: A juvenile specimen CUMZ 12182 from Tham Suea Lueang Temple, Loei and B paratype CUMZ 1562 from Tam Pha Bing Temple, Loei C-E Pollicaria mouhoti mouhoti C lectotype of Hybocystis mouhoti NHMUK 20130071 / 1 from Lao Mountains, Camboja D holotype of ' Pollicaria nicoarlingi ' MNHN-IM- 2000 - 37277, and E specimen CUMZ 12166 from Wang Daeng Cave, Phitsanulok F Pollicaria myersii, holotype of ' Pollicaria huberi ' NHMUK 20180253. Photo: F. Prugnaud, MNHN (D).

Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale.