Artema, Walckenaer, 1837

Aharon, Shlomi, Huber, Bernhard A. & Gavish-Regev, Efrat, 2017, Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae), European Journal of Taxonomy 376 (376), pp. 1-57 : 46

publication ID 10.5852/ejt.2017.376

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Artema View in CoL View at ENA sp. c from Sudan, Iran, Pakistan, and India

Figs 180–200 View Figs 180–185 View Figs 186–194 View Figs 195–200 , 207 View Figs 201–207 , 214 View Figs 208–214

Material examined

SUDAN: 8 ♂♂, 2 juvs, Khartoum [15.54° N, 32.53° E], 1960–1964, J.S. Cloudsley-Thompson leg. ( MRAC 121706 , 121718, 169130, 127506); GoogleMaps 1 ♂, in pure ethanol, Khartoum, 20 Aug. 2014, M. Siyam leg. ( ZFMK Siy 1); 1 ♂, 2 ♀♀, New Halfa [15.32° N, 35.60° E], 450 m a.s.l., Jun. 2013, M. Siyam leg. ( ZMB, 3 vials); GoogleMaps 1 ♀, in pure ethanol, same locality, 10 Aug. 2014, in house, M. Siyam leg. ( ZFMK Siy 2); 2 ♀♀, arrived in NY in engine parts from Sudan, Jun. 1995, K. Devage leg. ( AMNH).

IRAN: 1 ♀, Fars Prov., Rafsanjan, Mirza Cave (30.6° N, 56.3° E), 2300 m a.s.l., date not given, S.M. Tahami leg. ( ZMSU); GoogleMaps 1 ♀, Kerman Prov., Sirjan, Oota Cave (29.6° N, 55.1° E), 1880 m a.s.l., date not given, M.S. Tahami leg. ( ZMSU). GoogleMaps

PAKISTAN: 2 ♀♀, Baluchistan, Muslim Bagh Village (30.783° N, 67.766° E), 7 Sep. 2005, S.V. Ovchinnikov leg. ( ZMMU); GoogleMaps 1 ♂, Baluchistan, Quetta Town [30.18° N, 66.99° E], in house, 27 Jul. 2005, S.V. Ovchinnikov leg. ( MMU). GoogleMaps

INDIA: 1 ♂, in pure ethanol, Rajastan, Durjana (29.00° N, 74.82° E), 181 m, in steppe with sparse vegetation and single trees, at wall, 10 Mar. 2011, P. Jäger leg. ( SMF); GoogleMaps 1 ♀, Rajastan [25.73° N, 73.34° E], 275 m, hill S Pali, 8 Jan. 1962, Ross and Cavagnaro leg. ( CAS) GoogleMaps .


These specimens are similar to A. doriae and to A. transcaspica , but females differ by a very inflated and prominent pale median area of the epigynal plate (arrow in Fig. 200 View Figs 195–200 ); females also tend to have wider epigyna:>1.6 × as wide as long, versus <1.6 in A. doriae and A. transcaspica . Dark median sclerite varies from brown and long (1.0 × length of epigynal plate) to very dark and short (0.5 × length of epigynal plate) to totally absent. Males differ from A. transcaspica ( Figs 125–127 View Figs 121–127 ) by deeper dorsal notch on distal margin of procursus and from A. doriae ( Figs 94–96 View Figs 90–96 ) by shapes of processes a and c. Males also with more distinct and robust ridge above modified hairs in lateral view ( Figs 186–189 View Figs 186–194 , arrows in Figs 190–191 View Figs 186–194 ). Preliminary molecular data (16S and CO1 sequences) indicate that specimens from India and Sudan are indeed conspecific (identical sequences in both genes), but clearly different from A. nephilit sp. nov., A. bunkpurugu , A. doriae , and A. atlanta : P-distances to these four species range from 12.8 to 19.0% for CO1 and 14.6 to 20.0% for 16S (J. Eberle, A. Valdez-Mondragón, D. Dimitrov, B.A. Huber, unpublished data). However, as long as species limits between A. transcaspica and A. doriae are not convincingly solved, we feel that adding yet another name to this group potentially creates more problems than it solves.


Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"


Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet


USA, New York, New York, American Museum of Natural History


USA, California, San Francisco, California Academy of Sciences


Zoologisches Forschungsmuseum Alexander Koenig


Museum für Naturkunde Berlin (Zoological Collections)


American Museum of Natural History


Zoological Museum, Saratov State University


Zoological Museum, Moscow Lomonosov State University


Forschungsinstitut und Natur-Museum Senckenberg


California Academy of Sciences











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