Tomosvaryella

Ramos-Pastrana, Yardany & Rafael, José Albertino, 2021, Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species, Zootaxa 4985 (1), pp. 37-68 : 65

publication ID	https://doi.org/ 10.11646/zootaxa.4985.1.2
publication LSID	lsid:zoobank.org:pub:5B23C14A-C7AB-4F22-B0A8-79E700F29FFF
DOI	https://doi.org/10.5281/zenodo.5217542
persistent identifier	https://treatment.plazi.org/id/507C87C0-CD70-1C51-EEEA-3998FA52F89D
treatment provided by	Plazi
scientific name	Tomosvaryella
status

Treatment

Key to the Tomosvaryella species from Colombia

1 Hind trochanter with two bare spines ( Fig. 42 View FIGURES 37–44 ); phallus with central branch having three stout, reclined, apical spines and one of the lateral branches with one short reclined spine ( Fig. 50 View FIGURES 45–52 ).................................... T. martae View in CoL sp. nov.

- Hind trochanter with one distinct protuberance projected or slightly elevated ( Figs 2 View FIGURES 1–10 , 12 View FIGURES 11–20 , 26 View FIGURES 21–28 , 54 View FIGURES 53–62 , 64 View FIGURES 63–72 , 74 View FIGURES 73–82 , 84 View FIGURES 83–92 , 94 View FIGURES 93–102 , 104–108 View FIGURES 103–108 , 122 View FIGURES 120–123 ); phallus branches with variable reclined spines, if three spines, then spines medially placed on the branches ( Figs 8 View FIGURES 1–10 , 18 View FIGURES 11–20 , 34 View FIGURES 29–36 , 60 View FIGURES 53–62 , 70 View FIGURES 63–72 , 80 View FIGURES 73–82 , 90 View FIGURES 83–92 , 100 View FIGURES 93–102 , 114 View FIGURES 109–116 , 129 View FIGURES 124–131 ).............................................................................. 2

2 Hind trochanter slightly elevated, truncated, without acute protuberance ( Fig. 94 View FIGURES 93–102 ); syntergosternite 8 sub-hemispherical ( Fig. 96 View FIGURES 93–102 ); right surstylus wider than left, with distinct external submedian lobe ( Figs. 98–99 View FIGURES 93–102 ); apex of phallic guide thickened ( Fig. 100 View FIGURES 93–102 ); phallus with central branch stout, somewhat coiled ( Fig. 100 View FIGURES 93–102 ); ejaculatory apodeme fan-shaped ( Fig. 101 View FIGURES 93–102 ).................................................................................................. T. subvirescens View in CoL

- Hind trochanter with distinct protuberance ( Figs 2 View FIGURES 1–10 , 12 View FIGURES 11–20 , 26 View FIGURES 21–28 , 42 View FIGURES 37–44 , 54 View FIGURES 53–62 , 64 View FIGURES 63–72 , 74 View FIGURES 73–82 , 84 View FIGURES 83–92 , 104–108 View FIGURES 103–108 ), if no distinct protuberance, then with a small elevation having distinct ascendant and descendent margins with obtuse apex at higher point ( Fig. 122 View FIGURES 120–123 ); syntergosternite 8 somewhat directed to right side ( Figs 27–28 View FIGURES 21–28 , 43–44 View FIGURES 37–44 ); surstyli somewhat similar ( Figs 4–5 View FIGURES 1–10 , 14–15 View FIGURES 11–20 , 30–31 View FIGURES 29–36 , 46–47 View FIGURES 45–52 , 56–57 View FIGURES 53–62 , 66–67 View FIGURES 63–72 , 86–87 View FIGURES 83–92 , 110–111 View FIGURES 109–116 ); apex of phallic guide slender ( Figs 50 View FIGURES 45–52 , 60 View FIGURES 53–62 ); phallus branches with the same diameter, not coiled ( Figs 8 View FIGURES 1–10 , 18 View FIGURES 11–20 , 34 View FIGURES 29–36 , 50 View FIGURES 45–52 , 60 View FIGURES 53–62 , 70 View FIGURES 63–72 , 80 View FIGURES 73–82 , 90 View FIGURES 83–92 , 114 View FIGURES 109–116 , 129 View FIGURES 124–131 ); ejaculatory apodeme generally sandal-shaped ( Figs 9 View FIGURES 1–10 , 19 View FIGURES 11–20 , 35 View FIGURES 29–36 , 51 View FIGURES 45–52 , 61 View FIGURES 53–62 , 71 View FIGURES 63–72 , 91 View FIGURES 83–92 , 115 View FIGURES 109–116 , 130 View FIGURES 124–131 ) ................................................................................................... 3

3 Hind trochanter with bulging, drop-shaped protuberance (Fig. 184); both surstyli with external and internal margins somewhat straight ( Fig. 86–87 View FIGURES 83–92 )............................................................................. T. similis View in CoL

- Hind trochanter different ( Figs 2 View FIGURES 1–10 , 12 View FIGURES 11–20 , 26 View FIGURES 21–28 , 42 View FIGURES 37–44 , 54 View FIGURES 53–62 , 64 View FIGURES 63–72 , 74 View FIGURES 73–82 , 94 View FIGURES 93–102 , 104–108 View FIGURES 103–108 , 122 View FIGURES 120–123 ); both surstyli with external and internal margins sinuous ( Figs 4–5 View FIGURES 1–10 , 14–15 View FIGURES 11–20 , 66–67 View FIGURES 63–72 , 110–11 View FIGURES 109–116 , 125–126 View FIGURES 124–131 ), in case of doubts, apex distinctly hooked ( Fig. 30–31 View FIGURES 29–36 , 56–57 View FIGURES 53–62 )....... 4

4 Apex of phallic guide distinctly angled from base, without basal spine ( Figs 8 View FIGURES 1–10 , 129 View FIGURES 124–131 )................................ 5

- Apex of phallic guide lined with or slightly above the main body of the structure, with or without basal spine ( Figs 18 View FIGURES 11–20 , 34 View FIGURES 29–36 , 50 View FIGURES 45–52 , 60 View FIGURES 53–62 , 70 View FIGURES 63–72 , 80 View FIGURES 73–82 , 90 View FIGURES 83–92 , 114 View FIGURES 109–116 ).................................................................................... 6

5 Hind trochanter with short protruding protuberance with anterior margin somewhat straight and posterior margin descending, yellow-setose ( Fig. 2 View FIGURES 1–10 ); surstyli similar in shape ( Figs 4–5 View FIGURES 1–10 ); left surstylus with external margin sinuous; right surstylus somewhat straight dorsally in lateral view ( Figs 6–7 View FIGURES 1–10 ); phallus with no projection before trifurcation and two branches with small reclined spines ( Fig. 8 View FIGURES 1–10 )............................................................................ T. galapagensis View in CoL

- Hind trochanter with small triangular protuberance with obtuse apex at highest point, white-setose ( Fig. 122 View FIGURES 120–123 ); Surstyli asymmetrical, left surstylus with external margin somewhat straight ( Figs 125–126 View FIGURES 124–131 ), right surstylus dorsally distinctly lobate in lateral view ( Fig. 128 View FIGURES 124–131 ); phallus with distinct projection before trifurcation and two branches with stout reclined spines ( Fig. 129 View FIGURES 124–131 )........................................................................................ T. venezuelana View in CoL

6 Hind trochanter with protuberance somewhat squared, apex straight, yellow-setose apically ( Fig. 26 View FIGURES 21–28 ).................................................................................................... T. macarenensis View in CoL sp. nov.

- Hind trochanter with protuberance somewhat triangular having angled margins or rectangular with straight margins ( Figs 54 View FIGURES 53–62 , 64 View FIGURES 63–72 , 74 View FIGURES 73–82 , 104–108 View FIGURES 103–108 , 112 View FIGURES 109–116 ).................................................................................. 7

7 Hind trochanter with protuberance somewhat rectangular, with margins straight ( Figs 64 View FIGURES 63–72 , 104–108 View FIGURES 103–108 ).................... 8

- Hind trochanter with protuberance somewhat triangular with margins angled ( Figs 12 View FIGURES 11–20 , 54 View FIGURES 53–62 , 74 View FIGURES 73–82 )........................ 9

8 Surstyli setulose with high external lateral margins ( Fig. 66–67 View FIGURES 63–72 ); surstyli somewhat similar in shape, robust with apexes curved inward, inner and outer margins sinuous ( Fig. 66–67 View FIGURES 63–72 ); hind trochanter with long yellow-setulose protuberance at apex ( Fig. 64 View FIGURES 63–72 ); apex of phallic guide simple and conical, ventrally serrated ( Fig. 70 View FIGURES 63–72 ); central branch of phallus with three stout spines ( Fig. 70 View FIGURES 63–72 )........................................................................................ T. prostata View in CoL

- Surstyli bare, without high external lateral margins ( Figs 110–111 View FIGURES 109–116 ); surstyli similar in shape, robust with apexes curved inward, inner and outer margins sinuous ( Fig. 110–11 View FIGURES 109–116 ); hind trochanter with variable stout spiniform protuberance ( Figs 104–108 View FIGURES 103–108 ); apex of phallic guide short with a spine ( Fig. 114 View FIGURES 109–116 ); phallus with two branches bearing a fine spine ( Fig. 114 View FIGURES 109–116 )....... T. tuberculata

9 Hind trochanter with protuberance entirely white-setose ( Fig. 12 View FIGURES 11–20 ); surstylus elongated, somewhat tapered apically, both almost the same length, right surstylus slightly shorter ( Figs 14–15 View FIGURES 11–20 ), downward-directed in lateral view ( Figs 16–17 View FIGURES 11–20 ); apex of phallic guide short and with a fine spine ( Fig. 18 View FIGURES 11–20 ), one branch of phallus with a small spine ( Fig. 18 View FIGURES 11–20 )................... T. lynchi View in CoL

- Hind trochanter with protuberance with few spaced setae not entirely covered with white setae ( Fig. 54 View FIGURES 53–62 , 74 View FIGURES 73–82 ); surstylus variable as in Figs 57 View FIGURES 53–62 , 77 View FIGURES 73–82 ; apex of phallic guide without spine ( Fig. 80 View FIGURES 73–82 ), if spine present, then it is stout ( Fig. 60 View FIGURES 53–62 ); with central branch of phallus bearing stout spines ( Fig. 80 View FIGURES 73–82 ) or row of small spines ( Fig. 60 View FIGURES 53–62 )........................................... 10

10 Surstyli long, distinctly with hooked-tips, without setose basal lobe ( Fig. 56–57 View FIGURES 53–62 ); apex of phallic guide with stout spine ( Fig. 60 View FIGURES 53–62 ); phallus central branch with row of small spines ( Fig. 60 View FIGURES 53–62 ); ejaculatory apodeme sandal-shaped ( Fig. 61 View FIGURES 53–62 )..................................................................................................... T. mexicanensis View in CoL

- Surstyli robust, indistinctly with hooked-tips, with basal lobe setose ( Fig. 76–77 View FIGURES 73–82 ); apex of phallic guide without spine ( Fig. 80 View FIGURES 73–82 ); phallus with central branch with three distinct spines ( Fig. 80 View FIGURES 73–82 ); ejaculatory apodeme with apex truncated ( Fig. 81 View FIGURES 73–82 )................................................................................................... T. scopulata View in CoL