Tomosvaryella macarenensis, Ramos-Pastrana & Rafael, 2021

Ramos-Pastrana, Yardany & Rafael, José Albertino, 2021, Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species, Zootaxa 4985 (1), pp. 37-68 : 43-46

publication ID 10.11646/zootaxa.4985.1.2

publication LSID


persistent identifier

treatment provided by


scientific name

Tomosvaryella macarenensis

sp. nov.

Tomosvaryella macarenensis View in CoL sp. nov.

Figs 21–36 View FIGURES 21–28 View FIGURES 29–36 , 133 View FIGURE 133 A62E864A-73F9-46A5-BCF8-2DF274A08CC5

Diagnosis. Male. Hind trochanter with semi-transverse truncated long protuberance, apically with yellow setae. Surstyli subsymmetrical, slender, as long as epandrium, with hooked-tip, right surstylus slightly shorter, both surstyli somewhat straight in lateral view. Cerci elongated. Phallic guide short, with apex simple, bearing one small seta and tip angled at about 30 degrees. Phallus with one ejaculatory duct with two spines, one duct with a single spine and one duct without spine.

Description. MALE (holotype). Body length 3 mm; wing length 3 mm. Head ( Figs 21–22 View FIGURES 21–28 ). Holoptic, eyes contiguous for seven facets. F, EM, V (mm) = 0.2, 0.1, 0.1. Frons grayish-pruinose. Postcranium black, dorsally brownish-pruinose, ventrally grayish-pruinose. Antenna brown; pedicel with three dorsal and one ventral setae; postpedicel dorsally clearer, with apex filiform, LPP/WPP = 2.4; Labellum brown. Thorax ( Fig. 22–23 View FIGURES 21–28 ). Postpronotal lobe yellow. Scutum dark-brown, shiny, with dense brownish-pruinosity medially. Notopleuron grayish-pruinose. Scutellum dark-brown, grayish-pruinose, with inconspicuous band of setae. Mesopleuron light-brown. Mediotergite dark-brown to black, grayish-pruinose. Wing ( Fig. 24 View FIGURES 21–28 ). LW/MWW = 4. LTC/LFC = 0.5. Membrane hyaline, almost completely microtrichose except as follow: cell bc bare; cell c bare on proximal quarter; cell sc bare on proximal three quarters; cell r 1 bare on proximal quarter; cell bm bare on proximal half; cell cup bare on proximal anal quarter; lobe bare on distal three quarters. Halter with stem and knob yellow, base brown. Legs ( Fig. 25 View FIGURES 21–28 ). Coxae dark-brown; trochanters light-brown; femora dark-brown except for base and apex yellow; tibiae dark-brown, with base and apex yellow; tarsomeres 1-3 yellow, 4 and 5 brown. Hind trochanter with long semi-transverse, truncated, apically with yellow setae protuberance ( Fig. 25–26 View FIGURES 21–28 ). Abdomen ( Figs 22, 27–28 View FIGURES 21–28 ). Dark-brown to black, shiny, dorsally brownish pruinose. Tergite 1 with six lateral setae. Tergites and sternites 6 and 7 as in Fig. 29 View FIGURES 29–36 . Syntergosternite 8 brown, brownish-pruinose, shorter than tergite 5, with membranous area occupying less than half of syntergosternite in dorsal view ( Figs 27–28 View FIGURES 21–28 ). Terminalia ( Figs 29–36 View FIGURES 29–36 ). Epandrium brown and surstyli light brown. Surstyli ( Figs 30– 33 View FIGURES 29–36 ) subsymmetrical, slender, as long as epandrium, with hooked-tips; right surstylus slightly shorter, both surstyli somewhat straight in lateral view ( Figs 32–33 View FIGURES 29–36 ). Cerci elongated. Phallic guide short with apex simple, bearing one small reclinate seta and tip angled at about 30 degrees ( Fig. 34 View FIGURES 29–36 ). Phallus with one ejaculatory duct with two spines, one duct with a single spine and one duct without spine ( Fig. 34 View FIGURES 29–36 ). Ejaculatory apodeme somewhat sandal-shaped, darker near base ( Fig. 35 View FIGURES 29–36 ). Gonopods same length, left with acuter apex ( Fig. 36 View FIGURES 29–36 ).

FEMALE. Unknown.

Type material. HOLOTYPE ♂: COLOMBIA, Meta, PNN[Parque Nacional Natural] S[errania] Macarena, Caño Curia, Sendero Cachicamos , 3º21’N / 73º56’W, 460 m [eters], Malaise, 10.xi.–21.xii.2002, M. Duarte leg, “M2985” (1♂, IAvH) (photographed specimen) GoogleMaps ; PARATYPES ♂: “ idem, 09–24.ii.2003, W. Villalba leg, “M3524” (1♂, LEUA)”; “ idem, 17.i.–09.ii.2003, “M3529” (1♂, INPA)”. The holotype has the left hind leg glued to a triangle and pinned along specimen, left wing mounted in Canada balsam on a microslide with balsam, and terminalia placed in a microvial with glycerin, both pinned along the specimen. GoogleMaps

Geographical distribution. Colombia ( Fig. 133 View FIGURE 133 ).

Etymology. The specific epithet refers to the type locality, Macarena, National Natural Park, Colombia.

Habitat. The specimens were collected with Malaise traps at ground level at the Parque Nacional Natural Serrania La Macarena, within savanna vegetation and in the flooded forests of the Orinoco region of Colombia.

Taxonomic notes. Tomosvaryella macarenensis sp. nov. runs to T. longiseta Ale-Rocha 1996 in the couplet 13 of the key presented by Ale-Rocha & Souza (2011) based on hind trochanter with a short semi-transverse truncated protuberance. It differs from T. longiseta in having slender surstyli and apexes curved inward ( Figs 30–31 View FIGURES 29–36 ) (versus surstyli short with truncated apex in T. longiseta , figure 69 of Ale-Rocha 1996); apex of phallic guide short, with a small reclinate seta, phallus with two spines in central branch ( Fig. 34 View FIGURES 29–36 ) (versus apex of phallic guide long with stout projection, phallus with a fine spine in one branch, figure 73 of Ale-Rocha 1996).


Instituto Nacional de Pesquisas da Amazonia













GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF