Tobochares canaima, Girón & Short, 2021

Giron, Jennifer C. & Short, Andrew Edward Z., 2021, Review of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records, ZooKeys 1019, pp. 93-140 : 93

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Tobochares canaima

sp. nov.

Tobochares canaima sp. nov. Figs 3I View Figure 3 , 10D-F View Figure 10 , 11N View Figure 11 , 13 View Figure 13

Type material examined.

Holotype (male): "Venezuela: Bolívar: 5°51'N, 62°33'W; 1700 m; Auyan-tepui; Intercept trap; 7-14.ii.1994; leg. J.L. García, A. Chacón” (MIZA). Paratypes (7 exs.): Venezuela: Bolívar: Same data as holotype (6, MIZA, SEMC); " 5°46'50"N, 62°31'36"W; 2170 m; Auyan-tepui; yellow trap; 20.iv.1994; leg. L. Mesner, J.L. García” (1, MIZA).

Differential diagnosis.

Tobochares canaima can be recognized by its strongly convex body in lateral view, accompanied by elytral punctation uniform in size and degree of impression, with serial punctures seemingly aligned in rows, not impressed to form grooves; the interserial punctures are somewhat irregularly distributed in two or three rows (Fig. 10D, E View Figure 10 ). The general habitus and punctation of T. canaima are similar to those of T. arawak , T. kappel , and T. kolokoe . In T. kappel and T. kolokoe the interserial punctures form only one or two irregular rows (e.g., Fig. 3H View Figure 3 ). In T. arawak the pronotal and elytral punctation is shallower than in T. canaima (compare Fig. 10A View Figure 10 vs. Fig. 10B View Figure 10 ) and the apodemes of the median lobe are half the length of the median lobe in T. arawak (Fig. 11M View Figure 11 ), as opposed to one fourth of the length in T. canaima (Fig. 11N View Figure 11 ).


Size and form: Body length 1.9-2.1 mm. Body elongate oval, moderately convex (Fig. 10E View Figure 10 ). Color and punctation: Dorsal and ventral surfaces of body dark brown, with anterolateral margins of prothorax slightly paler (Fig. 10D, E View Figure 10 ); mouthparts yellow to orange; antennae light brown; legs reddish to dark brown, distally paler (orange), with paler (yellow) tarsi (Fig. 10F View Figure 10 ). Ground punctation on head, pronotum and elytra sharply marked; pronotal punctation dense (Fig. 10D View Figure 10 ). Head: Eyes in dorsal view with anterior margin oblique (anteriorly directed; e.g., Fig. 2D View Figure 2 ), and outer margins slightly bulging from outline of head; in lateral view, eyes not emarginate (e.g., Fig. 2E View Figure 2 ). Thorax: Elytra with serial punctures similar in size and degree of impression to interserial punctures, and only seemingly aligned longitudinally, not forming grooves; interserial punctures irregularly distributed in two or three rows (Fig. 10D View Figure 10 ). Metafemora mostly glabrous on anterior face (Fig. 10F View Figure 10 ). Elevation of mesoventrite forming a very low transverse carina (Fig. 10F View Figure 10 ). Metaventrite with distinct median, longitudinal, narrow glabrous area extending along posterior half (Fig. 10F View Figure 10 ). Abdomen. Abdominal ventrites uniformly and very densely pubescent (Fig. 10F View Figure 10 ). Aedeagus (Fig. 11N View Figure 11 ). Basal piece nearly 0.45 × the length of a paramere; greatest width of a paramere nearly 0.6 × greatest width of median lobe; outer margins of parameres straight to widely and uniformly convex; apex of paramere obliquely rounded; median lobe roughly sagittate, rounded at apex; gonopore situated nearly at midlength of median lobe.


Noun in apposition. Named after the Canaima National Park in Venezuela, where the type locality is situated.


This species is known from the famous Auyan-tepui, which is also home to Angel Falls, the highest waterfall in the world. Collected at elevations of 1700-2170 m, this species is one of the relatively few water beetle taxa known from the "Pantepui Province", which comprises areas of the Guiana Shield which are greater than 1500 m in elevation. See Fig. 13 View Figure 13 .

Life history.

The only known series was collected in a flight intercept trap and a yellow pan trap. Nothing further is known about the habitat or biology of this species.