Belisana tarang, Zhu & Li, 2021

Zhu, Wenhui & Li, Shuqiang, 2021, Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia, Zootaxa 4963 (1), pp. 115-137 : 130-136

publication ID	https://doi.org/ 10.11646/zootaxa.4963.1.5
publication LSID	lsid:zoobank.org:pub:6B4B8F71-7B08-4027-A76B-7EE8C987C9BD
DOI	https://doi.org/10.5281/zenodo.4697008
persistent identifier	https://treatment.plazi.org/id/4E3C87F3-FFCE-FF84-C6EC-FCC7FC34F871
treatment provided by	Plazi
scientific name	Belisana tarang
status	sp. nov.

Treatment

Belisana tarang View in CoL sp. nov.

Figs 11–12 View FIGURE 11 View FIGURE 12 , 13K–L View FIGURE 13 , 14I–J View FIGURE 14 , 15F View FIGURE 15

Type material. Holotype: Male ( IZCAS Ar41916), near entrance of Tarang Cave (0°12’39.18”N, 100°24’58.62”E, elevation 877 m), Bukittinggi, West Sumatra, Sumatra, Indonesia, 13 May 2014, Z. Yao leg. GoogleMaps Paratypes: 1 male ( IZCAS Ar41917) and 2 females ( IZCAS Ar41918–41919), same data as holotype GoogleMaps .

Etymology. The specific name refers to the type locality and is a noun in apposition.

Diagnosis. This species resembles B. sandakan Huber, 2005 (see Huber 2005: 59, figs 360–375) with the same shape of the bulbal apophyses ( Fig. 12D View FIGURE 12 ), but males can be distinguished by the presence of a nearly triangular subdisto-prolateral sclerite (arrow 1 in Figs 11C View FIGURE 11 , 13K View FIGURE 13 ) and triangular retrolateral membranous flap (arrow f in Figs 11D View FIGURE 11 , 13L View FIGURE 13 ) on the procursus ( Figs 11A–D View FIGURE 11 , 13K–L View FIGURE 13 ), and by having the cheliceral distal apophyses directed laterally (arrow da in Figs. 12C View FIGURE 12 , 15F View FIGURE 15 ; distal apophyses directed downward in B. sandakan ). Females can be distinguished by having the frontal “epigynal” pockets laterally on a sclerotized plate (arrow ep in Figs 12A–B View FIGURE 12 , 14I–J View FIGURE 14 ) and a long vulval anterior arch, 2/3 of the vulva length ( Figs 12B View FIGURE 12 , 14J View FIGURE 14 ) (vs. short vulval anterior arch, 1/2 of the vulva length). It can also distinguished by a longer total body length (total body length <1.4, “tiny” size in B. sandakan, Huber 2005: 3 ).

Description. Male (holotype, IZCAS Ar41916): Total length 1.46 (1.54 with clypeus), carapace 0.63 long, 0.63 wide, opisthosoma 0.91 long, 0.69 wide. Leg I missing (see variation), leg II: 7.45 (2.03 + 0.25 + 1.88 + 2.63 + 0.66), leg III: 3.23 (0.88 + 0.13 + 0.88 + 1.09 + 0.25), leg IV: 5.07 (1.47 + 0.19 + 1.19 + 1.63 + 0.59). Distance PME-PME 0.06, diameter PME 0.08, distance PME-ALE 0.08, AME absent. Habitus as in Figs 12E–F View FIGURE 12 . Carapace pale, with yellow median marks. Ocular area not elevated. Clypeus unmodified. Chelicerae ( Figs. 12C View FIGURE 12 , 15F View FIGURE 15 ) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.23). Sternum pale, approximately as wide as long (0.38). Thoracic furrow absent. Legs pale, without darker rings. Opisthosoma pale, without spots. Palps as in Figs 11A–B View FIGURE 11 ; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with small proximo-dorsal apophysis (arrow in Fig. 11B View FIGURE 11 ); procursus ( Figs 11A–D View FIGURE 11 , 13K–L View FIGURE 13 ) curved (arrow in Fig. 11A View FIGURE 11 ), complex distally, with large, nearly triangular subdisto-prolateral sclerite (arrow 1 in Figs 11C View FIGURE 11 , 13K View FIGURE 13 ), small subdisto-dorsal sclerite (arrow 2 in Figs 11C View FIGURE 11 , 13K View FIGURE 13 ), large distal membranous process (arrow in Figs 11D View FIGURE 11 , 13L View FIGURE 13 ) and triangular retrolateral membranous flap (arrow f in Figs 11D View FIGURE 11 , 13L View FIGURE 13 ); bulb ( Fig. 12D View FIGURE 12 ) with short, hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi, without spines and curved setae.

Female (IZCAS Ar41918): Similar to male, habitus as in Figs 12G–H View FIGURE 12 . Total length 1.39 (1.47 with clypeus), carapace 0.47 long, 0.47 wide, opisthosoma 1.00 long, 0.94 wide. Tibia I: 1.75; tibia I L/d: 29. Distance PME-PME 0.07, diameter PME 0.06, distance PME-ALE 0.07, AME absent. Sternum slightly wider than long (0.38/0.34). External female genitalia ( Figs 12A View FIGURE 12 , 14I View FIGURE 14 ) simple and flat, with pair of frontal “epigynal” pockets (arrow ep in Figs 12A–B View FIGURE 12 , 14I–J View FIGURE 14 ), 0.34 apart. Vulva ( Figs 12B View FIGURE 12 , 14J View FIGURE 14 ) with W-shaped anterior arch and pair of long, narrow pore plates (arrow pp in Figs 12B View FIGURE 12 , 14J View FIGURE 14 ), without serrated sclerites.

Variation: In male paratype (IZCAS Ar41917): Leg I: 10.57 (2.66 + 0.28 + 2.72 + 3.75 + 1.16); tibia I L/d: 45. Retrolateral trichobothria of tibia Iat 7% proximally; tarsus Iwith 16 distinct pseudopodomeres.

Natural history. The species was found in leaf litter and collected by small plot direct searching.

Distribution. Indonesia (Sumatra, type locality only; Fig. 16 View FIGURE 16 ).