Exocelina haia Shaverdo & Balke

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

19. Exocelina haia Shaverdo & Balke sp. nov. Figs 49 , 53

Type locality.

Papua New Guinea: Simbu Province, between Supa and Haia Villages (Airstrips), ca. 6°39'39.9"S, 145°00'28.4"E, 1032 m.

Type material.

Holotype: male "Papua New Guinea: Supa Haia, 1023m, 10.ix.2002, K. Sagata (WB1)" (ZSM). Paratypes: 2 males with the same label as the holotype (NHMW, ZSM). 1 male "Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500m, 12IX2002, Balke & Sagata, (PNG 005)" (ZSM).

Females of doubtful identity.

See under E. bacchusi .

Description.

Body size and form: Beetle small: TL-H 3.4-3.45 mm, TL 3.7-3.75 mm, MW 1.8 mm (holotype: TL-H 3.4 mm, TL 3.7 mm, MW 1.8 mm), with oblong-oval habitus.

Colouration: Dark brown, with paler sides of pronotum and head anteriorly. As in E. warasera sp. nov. ( Fig. 49 ).

Surface sculpture: Shiny dorsally, with extremely fine and sparse punctation and weakly impressed microreticulation. As in E. warasera sp. nov.

Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 very slightly concave.

Male: Protarsomere 4 with anterolateral seta thin and evenly curved, smaller than more laterally situated large seta. Protarsomere 5 ventrally with anterior band of ca. 80 and posterior row of two relatively long setae ( Fig. 53D ). Abdominal ventrite 6 with 6-8 lateral striae on each side. Median lobe short, curved, with broadly pointed apex in lateral view, and evenly tapering to pointed apex in ventral view ( Fig. 53A, B ). Paramere concave on dorsal side, with dorsal setae distinct: subdistal setae only slightly stronger and denser than proximal ( Fig. 53C ).

Female: Unknown.

Affinities.

Exocellina haia sp. nov. can be distinguished by the shape and setation of its median lobe and paramere and/or by its size and colouration from the species co-occurring in the same area ( E. bacchusi , E. craterensis Shaverdo & Balke, 2014, E. damantiensis , E. hintelmannae (Shaverdo, Sagata & Balke, 2005), E. warasera sp. nov.). For affinities within the group, see the “Key”.

Distribution.

Papua New Guinea: Simbu Province, Crater Mountain area ( Fig. 54 ). The species is named after Haia Village. The name is a noun in the nominative singular standing in apposition.

Figures

Figure 54. Map of the eastern part of New Guinea showing distribution of the species of the E. warasera group.

Figures 46 - 49. Habitus and colouration 46 Exocelina kobau sp. nov. 47 E. pulchella sp. nov. 48 E. warasera sp. nov. 49 E. haia sp. nov.

Figures 52, 53. 52 Exocelina warasera sp. nov. 53 E. haia sp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view.