Pinus sp.

Smith, MacKenzie A., Greenwalt, Dale E. & Manchester, Steven R., 2023, Diverse Fruits And Seeds Of The Mid-Eocene Kishenehn Formation, Northwestern Montana, Usa, And Their Implications For Biogeography, Fossil Imprint 79 (1), pp. 37-88 : 46

Pinus sp. S1

Text-fig. 3l, m

M a t e r i a l. USNM PAL 621980, 622300.

L o c a l i t y. Disbrow Creek.

D e s c r i p t i o n. Winged seed linear, incomplete; 10.4– 14.1 mm long, 4.4–5.0 mm wide at widest point; seed body on proximal side, obovate to elliptical, slightly detached from wing, distally rounded 2.2–3.2 mm long and 1.3– 1.5 mm wide; admedial edge of wing begins a third of the way up the seed body and rapidly becomes larger at the end of the seed body’s distal end; medial edge of wing straight to slightly concave; striations on wing slightly undulating, striations of similar thickness.

R e m a r k s. Pinus subg. Strobus (also known as

Haploxylon , the soft pines or white pines) have seed bodies that can disarticulate, such that the fossil seeds of this subgenus are often missing the seed body. This fossil falls into Wolfe and Schorn’s (1990) circumscription of atypical Pinus 4.

Figures

Text-fig. 3. Pinaceae from the Kishenehn Formation. a: USNM PAL 621424 Abies sp. seed. Scale bar 5mm. b: USNM PAL 621991 Larix sp. winged seed. Scale bar 2mm. c: USNM PAL 622093cf. Larix sp. ovuliferous cone. Scale bar 3mm. d: USNM PAL 776558 Picea sp. 1 winged seed. Scale bar 3mm. e: USNM PAL 776565 Picea sp. 2 seed. Scale bar 5mm. f: USNM PAL 619964 Pinaceae Cone 1. Scale bar 1cm. g: USNM PAL 776557 Pinaceae Cone 2. Scale bar 1cm. h: USNM PAL 620025 Pinaceae Cone 3. Scale bar 1cm. i: USNM PAL 776556 Pinus sp. C1. Scale bar 2cm. j: DMNH EPI. 48178 Pinus sp. L1 3-leaf fascicle. Scale bar 2cm. k: USNM PAL 776570 Pinus sp. C2 Pinus subg. Strobus female cone. Scale bar 1cm. l: USNM PAL 621980 Pinus subg. Strobus, Pinus sp. S1 winged seed. Scale bar 3mm. m: USNM PAL 622300 Pinus subg. Strobus, Pinus sp. S1 winged seed. Scale bar 5mm. n: USNM PAL 776567 Pinus subg. Strobus, Pinus sp. S2 seed. Scale bar 2mm. o: USNM PAL 620785 Pinus subg. Strobus, Pinus sp. S3 winged seed, with seed body disarticulated. Scale bar 2mm. p: USNM PAL 626064 Pinus subg. Pinus female cone scale Pinus sp. C3. Scale