Archisargidae Rohdendorf, 1962, 2012

Zhang, Junfeng, 2012, Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China, ZooKeys 238, pp. 57-76 : 59-61

Family Archisargidae Rohdendorf, 1962

Uranorhagionidae KY Zhang, Yang et Ren, 2010, p. 564, syn. n.

Origoasilidae KY Zhang, Yang et Ren, 2011, p. 995, syn. n.

Type genus.

Archisargus Rohdendorf, 1938

Included subfamilies.

Archisarginae Rohdendorf, 1962 and Uranorhagioninae KY Zhang, Yang et Ren, 2010, stat. n. (= Mostovskisarginae JF Zhang, 2010, syn. n.).

Redefinition.

Moderate- to large-sized flies. Body robust but usually narrow and long, strongly pubescent but devoid of bristles; first flagellomere of antenna unsegmented, arista well developed at tip of first flagellomere; hind legs stout and long, femora clavate; tibial spurs and pulvilliform empodium well developed; wing narrow and long, subpetiolate, alula absent; all longitudinal veins well developed, ending at wing margin; C running around entire wing margin although thinned near to, or beyond, wing tip, C and R strong, Sc and R1 long, R4+5 bifurcated, R2+3 usually straight and long but in some specific members R2+3 short and significantly curved [see Figure 4 , Daohugosargus eximius (KY Zhang et al., 2008) comb. n., originally Sharasargus eximius KY Zhang et al., 2008], in most representatives crossvein r-m meeting R4+5 distad to Rs fork but in some specific members far basad to Rs fork (see Figure 4 , Daohugosargus eximius ), origin of Rs usually basad to, but in some specific members distad to, d base (see Figure 4 , Daohugosargus eximius ), discoidal cell shifted distally of wing midpoint.

Remarks.

Rohdendorf (1962) defined the family Archisargidae based on a single poorly preserved wing from the Callovian-Oxfordian Karabastau Formation. Kovalev (1981) argued that this family was described from very poor material and thus nothing definite can be said about its systematic position. However, the type genus Archisargus Rohdendorf, 1938, the only representative of the Archisargidae , clearly has little in common with the Jurassic Rhagionidae : it is a large fly with a long (16 mm) narrow wing. Mostovski (1996a, 1996b) described many new species referred, respectively, to some new genera or the known genera and assigned these to Archisargidae . Immediately after these, he re-described the type species of type genus based on the holotype; and a redefinition of the Archisargidae was proposed: wing venation not, or slightly, costalized; C running around entire wing margin although thinned beyond wing tip or R4 end; R1 long; R4+5 bifurcated; crossvein r-m meeting R4+5 distad to Rs fork; base of discoidal cell distad to origin of Rs from R; M4, if present, connecting with discoidal cell ( Mostovski 1997).

Recently, numerous well preserved archisargid flies have been recovered from the Daohugou biota, China ( JF Zhang and HC Zhang 2003; JF Zhang 2010a, 2012b; KY Zhang et al. 2007a, 2007b, 2008a, 2009, 2010a, 2010b). The familial diagnosis may be further supplemented based on information derived from these new results.

Figures

Figure 4. Daohugosargus eximius (KY Zhang, Yang et Ren, 2008) comb. n. (originally Sharasargus eximius KY Zhang, Yang et Ren, 2008; after KY Zhang et al. 2008).