Mitella stylosa

M. Wakabayashi, 2006, Tiarella and Mitella, Flora of Japan, Volume 2 b: Angiospermae-Dicotyledoneae: Archichlamydeae (b), Kodansha, pp. 70-75 : 73-74

var. subramosa Wakab.

in Acta Phytotax. Geobot. 28: 120 (1977).

Mitella stylosa auct. non H.Boissieu: Ohwi in Acta Phytotax. Geobot. 1: 63 (1932), pro parte majore.

Japanese name: Charumeru-so.

Rhizomes rather short, thickened. Radical leaf blade widely ovate, deeply cordate, obtuse to subacute, 1.5-12 cm long, 1.4-8.5 cm wide; petiole 1-15 cm long. Flowering stems 12-50 cm tall. Inflorescences racemes, many-flowered. Flowers April to May. Calyx greenish or pale brownish or purplish; calyx tube broadly obconical to campanulate-obconical; calyx lobes erect or slightly recurved at tip, 0.9-1.5 mm long, triangular or ovate or narrowly ovate, acute, glandular along margin or entire. Petals generally 3-5-pinnatiparted; pinnae obliquely ascending, glandular dotted, purplish, 4-4.4 mm long. Stamens 0.9-1.7 mm long; anthers rarely with 1 or 2 tubercles at apex, ca. 0.7-1.3 mm long at anthesis. Disc prominent or inconspicuous. Styles 0.3-1.5 mm long; stigmas thickened, not lobed or shallowly or distinctly 2-or 4-lobed.

Chromosome number: 2n = 28 (Wakabayashi 1973).

Japan: W. Honshu (westward from Ibuki and Suzuka Mountains), Shikoku (rare in Ehime Pref.) and N. Kyushu. Moist, shady places along streams in mountainous areas. Endemic.

Icones: Kitamura & Murata, Herb. PI. 2: t. 35 299 and f. 72 3, as M. stylosa-, Wakabayashi, Acta Phytotax. Geobot. 28: 120, f. 8 (1977); Satake et al., Herb. PI. 2: photo. 153 1 & 2.