Euwallacea Hopkins

Hulcr, Jiri, 2010, New genera of Palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae) based on congruence between morphological and molecular characters, Zootaxa 2717, pp. 1-33 : 16

Genus Euwallacea Hopkins

Type species. Xyleborus wallacei (Blandford) , desig. Hopkins, 1915

Specimens examined. E. wallacei : Malaysia (holotype, BMNH); New Guinea, Madang Province, Ohu (BHJ); New Guinea, Morobe, Wau (4, Hulcr det. BBM); New Guinea (1, Hulcr det. FICB); New Guinea, Madang Province, Baiteta (1, Hulcr det. IRSNB); New Guinea, Gulf Province, Ivimka (1, Hulcr det. UCD); New Guinea, Madang Prov. (46, Hulcr det., MSUC), Oro Prov. (3, Hulcr det., MSUC), West Sepik (8, Hulcr det., MSUC). E destruens : Indonesia, Java, Gilolo (syntypes, 2 BMNH); New Guinea, Morobe Province, Mt. Kaindi, 2400m a.s.l. (1, Hulcr det. BBM); New Guinea, New Britain (5, BBM); New Guinea, Madang Province, Baiteta ((1, Hulcr det. IRSNB); PNG, Chimbu Prov. (15, Hulcr det., MSUC), Madang Prov. (100, Hulcr det., MSUC), West Sepik (9, Hulcr det., MSUC).

Diagnosis. Euwallacea is similar, and probably a sister or paraphyletic genus, to Wallacellus gen. n. and Fortiborus gen. n. ( Fig. 9 ). Characters distinguishing Euwallacea from Wallacellus include: upper part of eyes as large as lower part (in E. wallacei ), first segment of funicle slender, stalk-like, as long or longer than pedicel (except this is also present in W. striatulus ); pronotum quadrate to subquadrate (partially overlapping with Wallacellus which has subquadrate to rounded pronotum); protibiae triangular, with 6 or fewer denticles, protibial spur present (a single denticle separated from other equidistant denticles); sockets of tibial denticles enlarged. Characters distinguishing Euwallacea from Fortiborus gen. n. include: first segment of antennal club concave, circular around the club (not recurved, convex); antennal club rounded or taller than wide (antennal club broader than long in several Fortiborus ); anterior pronotal edge not elevated, lacking a row of serrations; protibiae triangular, with 6 or fewer denticles, protibial spur present. Characters distinguishing Euwallacea from Planiculus gen. n. include: Much larger size, subquadrate pronotum, pronotal psoterocoxal process slender and inconspicuous.

Comments. Euwallacea was insufficiently described in the original description ( Hopkins, 1915), and was later used as a vaguely defined unnatural group defined only by a broad declivity. We attempted to partially remedy the situation by transferring several species into new and hopefully monophyletic genera Fortiborus , Planiculus , and Wallacellus .

Figures

FIGURE 9. Morphology-based cladistic analysis of species previously placed in Euwallacea, Xyleborus, and several similar species of unclear affiliation. Combinable-component consensus of 72 MP trees, (113 steps). Numbers at nodes indicate Bremer support.