Embates cordiger Prena, 2005

Prena, Jens, 2005, The Middle American species of Embates Chevrolat (Coleoptera: Curculionidae: Baridinae), Zootaxa 1100 (1), pp. 1-151 : 1-151

39. Embates cordiger Prena sp. n.

(Fig. 146–148, 258)

Ambates sp. 12. Marquis 1991: 200

Holotype male (dissected), Costa Rica, labeled: “Sector Cocori, Fca/ de E. Rojas, 150m, Prov./ Limon, COSTA RICA, E/ Rojas , Nov 1991,/ L­N­286000, 567500”, CRI000 566428 ( INBC).

Paratypes 79 (16 males, 26 females, 37 not sexed), Costa Rica, labeled: same collecting site as holotype (with variants in wording) with following dates and barcodes : Jul 1991, CRI000 597164; Ago 1991: CRI000 582009, CRI000 582022; Nov 1991: CRI000 503445, CRI000 525124, CRI000 525126, CRI000 503442, CRI000 503446, CRI000 566701, CRI000 566703; Ene 1992, CRI000 333405; Mar 1992, CRI000 364783; 26.3– 24.4.1992, CRI000 767366; 26.6.– 16.7.1992, CRI000 702789; 10.– 30.9.1992, CRI000 989589; 9.– 30.11.1992, CRI000 862369; Ene 1994: CRI001 855750, CRI001 855796; Feb 1994: CRI001 846605–07; Mar 1994, CRI001 740178 ( INBC 23 ) ; “ COSTA RICA / Pr. Heredia / Puerto Viejo / Finca La Selva ”, “ R.J. Marquis coll./ No. 1056 28­VI 1982 ”, “ Piper / cenocladum”; same label except: No. 931 , 2­V 1982; No. 988, 29­V 1982; No. 1068, 4­VII 1982; No. 1033, 20­VII 1982; No. 984, 30­V 1982 (4 x); No. 237, 9­IV 1981, P. arieianum ( NMNH 10 ) ; “ COSTA RICA, Pr. Heredia / Pto. Viejo­La Selva B.S./ No. 266 22­III 1981 / R.J. Marquis coll.”, “ Piper / cenocladum” ( NMNH) ; “ Costa Rica, HEREDIA: 3/ km S Puerto Viejo, Est. / Biol. La Selva , 100 m,/ 19.– 22.2.2000, leg. Prena ” ( CWOB 2 , JPPC 7 ) ; “ Est. Pitilla, 700m, 9km S/ Sta. Cecilia, P.N. Guana­/ caste, Prov. Guan , COSTA/ RICA, C. Moraga ,/ Set 1991 / L­N­330200, 380200”, CRI000 519646 ( INBC) ; “ Costa Rica, Pr. Guanacaste / Est. Pitilla , 9km S Sta. Cecilia / 700m, P.N. Guanacaste / 6– 12.3.1996 leg. Prena ” ( JPPC) ; “Rio Sardinas, R.N. F.S. Barra del Colorado,/ Prov. Limón, COSTA RICA. 10 m. 1–14/ Feb 1994, F.V. Araya, L N/ 291500_564700 #2607”, CRI001 876874 ( INBC) ; “ Rio San Lorenzo, 1050m,/ Tierras Morenas, Z.P./ Tenorio, Prov. Guanacaste / Costa Rica, C. Alvarado / 10–20 feb 1992 / L­N 287800, 427600”, CRI000 768255 ( INBC) ; “ COSTA RICA: Prov. Heredia:/ 11km ESE La Virgen, 250–/ 350m, 10°21'N 84°03'W / leg. J. Prena, 6.– 11.4.2004 / INBio­OET­ALAS transect” ( INBC) ; “ COSTA RICA: Prov. Heredia:/ 10km SE La Virgen, 450–/ 550m, 10°20'N 84°05'W / 17.– 23.2.2003 / INBio­OET­ALAS transect”, “handcollecting/ leg. Jens Prena ”, INB0003229770–72 ( JPPC 3 ) ; same label except 11.– 16.3.2003, INB0003229902–06 ( INBC 5 ) , 17.– 20.3.2003, INB0003229979–81 ( INBC 3 ) , 8.– 13.4.2003, INB0003230045–56 ( JPPC 12 ) , 14.– 20.4.2003, INB0003230152, INB0003230156–60

( INBC 6); “ COSTA RICA: Prov. Heredia:/ 12 km SE La Virgen, 550–/ 650m, 10°19'N 84°05'W / Prena , 9.II.2001 / INBio­OET­ALAS transect”, INB0003209979 ( INBC); “ Alajuela / Upala, Faldas N./ E.U. Tenorio / 750 m, 19 Ab 1988/ Gonzalez,/ Soto” ( MUCR). Description. Habitus: total length 4.0– 5.8 mm (m=4.7, n=42). Color: integument rufous­castaneous; basic vestiture moderately dense, cupreous, dorsolateral pronotal vitta of cupreous scales moderately broad, ill­defined, scales velvety black in heart­shaped elytral macula (Fig. 146); venter with white scales on prosternum and along flank. Head: frontal fovea absent, rostrum rather slender, curved, subcylindrical (as Fig. 144), apex slightly expanded laterally, subcostate dorsolaterally but not medially, basolateral margin slightly edged, length of rostrum ♂♂ 1.04–1.15 x (m=1.08, n=16), ♀♀ 1.00–1.20 x (m=1.11, n=26) pronotal length, length of ante­antennal portion ♂♂ 0.36–0.39 x (m=0.38, n=16), ♀♀ 0.39–0.44 x (m=0.41, n=26) total rostral length, dorsal margin of antennal scrobe reaching rostral base before eye; length of funicular segments 1 and 2 subequal, club oblong ovate. Pronotum: length 0.95–1.02 x (m=0.98, n=42) maximum width, sides subparallel to slightly rounded in basal half, anterior portion tubulate; disk densely punctate, intervals granulose, subcostate dorsomedially or not. Elytra: length 1.65–1.82 x (m=1.73, n=41) width at humeri, width 1.26–1.38 x (m=1.33, n=41) maximum pronotal width, sides subparallel in basal half, apices rounded conjointly, preapical callus developed, striae fine, punctures indistinct, interstriae flat, 9 costate, 7 subcostate at subapical callus. Legs: slender, tibia nearly straight and parallel­sided, ventral margin with distal cluster of cupreous hairs, claws separate at base. Male: aedeagus as E. euchasma , with apex of aedeagus round (Fig. 147) .

Plant association. Piper cenocladum (Marquis 10, Prena 27), P. arieianum (Marquis 1; association probably accidental).

Distribution. Costa Rica, northern lowlands, Atlantic side (Fig. 258).

Specific epithet. The name is a compound Latin adjective derived from cor (heart) and gero (to exhibit).

Discussion. Embates cordiger belongs to the E. euchasma complex. The species is restricted to the Atlantic lowlands of Costa Rica (and probably Nicaragua), and can be distinguished from sympatrically occurring E. euchasma by the heart­shaped elytral macula, separated tarsal claws and smaller size. They are associated with different plants.