Embates rutilus Prena, 2005

Prena, Jens, 2005, The Middle American species of Embates Chevrolat (Coleoptera: Curculionidae: Baridinae), Zootaxa 1100 (1), pp. 1-151 : 1-151

52. Embates rutilus Prena sp. n.

(Fig. 174, 248)

Ambates sp. 7 . Marquis 1991: 200

Holotype male (dissected), Costa Rica, labeled: “Est. Pitilla, 700 m, 9 km S Sta. / Cecilia, P.N. Guanacaste, Prov./ Guan., COSTA RICA. Feb a mar/ 1993. P. Ríos. L­N­330200, 380200”, CRI001 188027 ( INBC).

Paratypes 57 (29 males, 28 females), Costa Rica and Panama, labeled: “Sect. San Ramon de Dos Rios, Prov./ Alaju, COSTA RICA, 620m, 25 MAR­12/ ABR 1995. C. Cano./ L N 318100 381900”, CRI002 204767 ( INBC); “Est. Pitilla. 9 km S. Santa Cecilia, P.N./ Guanacaste, Prov. Guana, COSTA RICA./ 700 m. May 1994, C. Moraga, L N/ 330200_380200 #2999”, CRI001 906787 ( INBC); “Est. Pitilla, 700m, 9 km S Sta./ Cecilia, P.N. Guanacaste, Prov./ Guana., COSTA RICA. 18 abr a 9/ may 1993. C. Moraga./ L­N­330200 380200”, CRI001 308249 ( INBC); “Est. Pitilla, 700m, 9 km S/ Sta Cecilia, Prov. Guana./ COSTA RICA. C. Moraga/ Abr 1991./ L­N­330200 380200”, CRI000 687160 ( INBC); “Estacion Pitilla 9 km S. de Santa/ Cecilia, Prov. Guana, COSTA RICA./ 700 m. JUL 1995. C. Moraga./ L N 329950 380450 #5352”, CRI002 173129 ( INBC); “Estac. Pitilla, 700m, 9 km S/ Santa Cecilia, Guanac. Pr/ COSTA RICA. Oct 1989 / C. Moraga & P. Rios/ 330200, 380200”, CRI000 131966 ( INBC); “Est. Pitilla, 700m. 9 km S/ Santa Cecilia, P.N. Guana­/ caste, Prov. Guan. COSTA/ RICA, P. Rios, Set 1991,/ L­N­ 330200, 380200”, CRI000 610372 ( INBC); “Rio San Lorenzo, 1050m,/ Tierras Morenas, Z.P./ Tenorio, Prov. Guanacaste / Costa Rica, Abr 1992 / F. Quesada/ L­N 287800,427600”, CRI000 868149 ( INBC); “Rio Sn Lorencito, 900m / Res. For. Sn Ramon, 5 km N/ Col. Palmarena, Alajuela / COSTA RICA. Mar 1990 / Curso Carabidae / 244500­470700”, CRI000 158221 ( INBC); “R. San Lorencito, 900 m, R. F. San/ Ramón, 5 km N de Colonia/ Palmareña, Prov. Alaju., COSTA/ RICA. 13–18 Jun 1993. I Curso/ Scarabaeidae . L­N­ 244500, 470700”, CRI001 365076 ( INBC); “ Alajuela / Colonia Libertad,/ Upala (Bosque)/ 450 m, 6­May 1988 / H. Lezama” ( MUCR); “ Guanacaste / Parque Nat. Rincon/ de la Vieja, Liberia / 810 m, 22­Nov 1987 / H. Lezama” ( MUCR); “ COSTA RICA Prov. Limón, No/ protegida. Hitoy Cerere, Send. Hacia/ Rio Moín. 400 m. 28 SEP 2000. W./ Arana./ L_S_400350_573200 #63419” ( INBC); “ COSTA RICA, Pr. Heredia / Pto. Viejo­La Selva B.S./ No. 83 31­VIII 1980 / R.J. Marquis coll.”, “ Piper / cenocladum”; same label except: No. 234 9­III 1981; No. 678 30­VI 1979; No. 222 6­VII 1981, Piper imperiale ; No. 568 15­X­1981, Piper imperiale ; No. 752 I­10­1982, Piper biseriatum ; No. 660 8­XI 1981, Piper biseriatum ; No. 443 VI­1­1981, Piper biseriatum ; No. 365 1­V­1981, Piper biseriatum ( NMNH 10); “ COSTA RICA / Pr. Heredia / Puerto Viejo/ Finca La Selva”, “R.J. Marquis coll./ No. 649 1­XI 1981 ”, “ P. melanocladum ”; same label except: No. 453, 1­VI 1981; No. 374, 5­V 1981; No. 281, 1­IV 1981; No. 279, 29­III 1981; No. 106, 6­X 1980; No. 57, 30­VII 1980; No. 1321, 5­IX 1979; No. 847, 16­II 1982; No. 1296, 27­IX 1983, Piper #11; No. 1107, 10­VIII 1982, Piper #3; No. 386, 6­V 1981, P. imperiale ; No. 935, 28­IV 1982, P. biseriatum ; No. 626, 28­X 1981, P. biseriatum ; No. 84, 7­II 1982, P. obliquum ; No. 637, 29­X 1981, P. arboreum ( NMNH 16); “ COSTA RICA, Heredia:/ Est. Bio. La Selva, 50–/ 150m, 10°26'N 84°01'W / Sept 1992, INBio­OET”, CRI001 216979; same label except Aug 1994, CRI001 245333 ( INBC 2); “ COSTA RICA, Heredia: La Selva Biol. Sta./ 3 km S Pto. Viejo, 100m / 20°26'N 84°01'W ”, “ 21.IV.2001 / leg. J. Prena”; same label except 19­ 22.2.2000; 20.IV.2001 (2 x); 22.IV.2001; 16.III.2001; 18.III.2001 ( CHAH, JPPC 6); “ COSTA RICA, Prov. Heredia:/ 10km SE La Virgen, 450–/ 550m, 10°20'N 84°05'W / 1.3.2000 / INBio­OET­ALAS transect” ( JPPC), 17.– 23.2.2003, INB0003229806 ( JPPC), 11.– 16.3.2003, INB0003229876–77 ( INBC 2), 14.– 20.4.2003, INB0003230165 ( JPPC); “Tucurrique/ Costa Rica ”, “Coll Schild/ & Burgdorf” ( NMNH); “ PANAMA: Chiriqui Pr./ Reserva la Fortuna/ Cont. Divide Trail/ 17­VI­1994, 1200m./ A.R. Gillogly” ( HPSC); “ PANAMA, Chiriquí: Res./ La Fortuna, 1100 m./ 8° 44'N 82° 14'W, 21.– 25.3.2001, lg. Prena” ( JPPC); “ PANAMA: Chiriqui / Continental divide trail/ 17 May 1996 / R. Turnbow” ( CMNC).

Description. Habitus: as Fig. 170, total length 6.3–8.2 mm (m=7.3, n=51). Color: integument rufous­castaneous; basic vestiture of small to medium­sized cupreous scales, scales yellow in broad dorsolateral pronotal and fuzzy elytral vittae between humerus and preapical callus (Fig. 174); venter with yellow scales on prosternum and along flank. Head: frontal fovea minute or obsolete, rostrum moderate, subcylindrical (as Fig. 171), sides attenuated between apex and antennal insertion, costate dorsomedially, basolateral margin moderately edged, length of rostrum ♂♂ 1.15–1.38 x (m=1.23, n=26), ♀♀ 1.20– 1.41 x (m=1.29, n=25) pronotal length, length of ante­antennal portion ♂♂ 0.33–0.39 x

(m=0.36, n=26), ♀♀ 0.40–0.46 x (m=0.42, n=25) total rostral length, dorsal margin of antennal scrobe reaching rostral base before eye; funicular segment 2 longer than 1, club oblong ovate. Pronotum: length 0.82–0.95 x (m=0.89, n=51) maximum width, sides subparallel to slightly rounded in basal half, then roundly narrowed, anterior portion tubulate; disk densely and very shallowly punctate, intervals very narrow, subcostate dorsomedially or not. Elytra: length 1.75–1.98 x (m=1.87, n=48) width at humeri, width 1.26–1.42 x (m=1.32, n=51) maximum pronotal width, sides subparallel in basal half, apices rounded separately, preapical callus developed moderately, striae fine, punctures indistinct, interstriae flat, 9 subcostate. Legs: moderately stout, tibia slightly curved, ventral margin with distal cluster of cupreous hairs, tarsal claws flat and approximate at base. Male: aedeagus as E. consimilis .

Plant association. Several species of Piper with the shoot­apex emerging from within the leaf­base at flowering nodes: P. biseriatum (Marquis 7, Prena 4), P. melanocladum (Marquis 9, Prena 2), P. cenocladum (Marquis 3, Prena 3, Letourneau 1, Lezama 1), P. imperiale (Marquis 3), P. euryphyllum (Marquis 1, Prena 1), P. arboreum (Marquis 1), P. hastigerum (Prena 1), P. obliquum (Marquis 1).

Rearing records. Piper cenocladum (Letourneau 1).

Distribution. Costa Rica and Panama, Atlantic side of Cordilleras (Fig. 248).

Specific epithet. The name is a Latin adjective meaning “reddish”.

Discussion. Embates rutilus belongs to a group of species with singularly rounded elytral apices, represented in Middle America by E. justini , E. rhombifer and E. consimilis . The elytral vitta is ill­defined in the specimens from the lowlands, but well­limited in the specimens from La Fortuna, at higher elevations.