Ganesella capitium (Benson, 1848)

Sutcharit, Chirasak, Backeljau, Thierry & Panha, Somsak, 2019, Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae), ZooKeys 870, pp. 51-76 : 51

Ganesella capitium (Benson, 1848) Figs 2A, B , 8

Helix capitium Benson, 1848: 160. Hanley and Theobald 1870: 7, pl. 14, fig. 5. Tryon 1887: 74, pl. 14, fig. 99.

Helix (Planispira) capitium : Nevill 1878: 78.

Trochomorpha capitium : Morlet 1889: 124, 125.

Ganesella capitium : Pilsbry 1895: 170, pl. 55, fig. 18. Blanford 1903: 278. Gude 1914: 196, 197. Zilch 1960: 610, fig. 2140. Zilch 1966: 202. Richardson 1985: 132, 133.

Darwininitium shiwalikianum Budha & Mordan in Budha et al. 2012: 21-23, figs 2-4. Type locality: Kasara near Tamor Lake, Chitwan National Park, Central Nepal. New synonym.

Type locality.

Sicrigali province Bahar Indiae Orientalis [Bihar State, India].

Material examined.

Type specimens. Three syntypes of Helix capitium are in Benson’s collection. The specimen that closely matched with the measurement in the original description is designated here as the lectotype UMZC I.102385/1 ( Fig. 2A , height 13.3 mm, width 13.5 mm), and the other two as paralectotypes UMZC I.102385/2-3 (2 shells; Fig. 2B , height 14.4 mm, width 13.8 mm).

Additional material.

Bahar Province, India: ZMB ex. Albers coll. 1 lot (1 shell). South India: NHMUK ex. Godwin-Austen coll. no. 501 (2 shells).

Remarks.

Budha et al. (2012) described Darwininitium shiwalikianum from Chitwan National Park, Nepal. The holotype has a trochoid, brownish shell with irregular opaque white spots. As such, it appears to be identical to the type specimens of Ganesella capitium . In addition, the type localities of D. shiwalikianum and G. capitium are geographically quite close to each other. Hence, in line with our earlier conclusions about the genus Darwininitium , we tentatively regard D. shiwalikianum as a junior subjective synonym of G. capitium .

Shell and genitalia have been described in detail by Budha et al. (2012). The unique and distinctive characters of G. capitium are its small, relatively elevated, trochoid shell. Aperture open sublaterally. Whorls slightly convex with wide and shallow suture. Last whorl angular with weak peripheral keel. Shell colour brownish with whitish-opaque, irregular, spots or streaks. Genitalia with short atrium; cylindrical, short penis, about half as long as the vagina, and proximally with blackish, spongy tissue. Epiphallus and flagellum together short, about as long as the penis, but flagellum longer than epiphallus. Internal wall of penis with numerous longitudinal pilasters.

Figures

Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133.

Figure 8. Geographic distribution of Ganesella spp. and Globotrochus spp.