Inella harryleei, Rolan & Fernandez-Garces, 2008,

Fernandes, Maurício Romulo & Pimenta, Alexandre Dias, 2019, Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil, European Journal of Taxonomy 517, pp. 1-60: 46

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Inella harryleei


“Inella” harryleei Rolán & Fernández-Garcés, 2008 

Figs 2LView Fig, 30–32View FigView FigView Fig

Material examined USA – Florida State • [1, d] spec.; off Saint Petersburg ; 28°26´56″ N, 84°40´37″ W; 63 m depth;

24 May 2012; G. Paulay leg.; FLMNH 450495.


Description of basic anatomy

OPERCULUM. Yellowish, ovate, flat, moderately thin but rigid, semi-transparent, 3.25 distinct whorls, nucleus slightly eccentric, dislocated ~37% from center toward margin; diameter of operculum does not exceed diameter of opercular pouch.

JAW. Wing-shaped; outer side with ~30 rows of scales, each scale often rhombus-shaped/squared, occasionally puzzle-shaped, rectangular or irregular; micro-pores in the posterior region, up to 380 nm in diameter; inner side with scales gem-like or leaf-shaped, surface smooth; scales of outer side 10.9– 16.5 µm long, 9.1–11.0 µm wide, ratio length/width 1.0–1.6 (rhombus-shaped/squared), 7.9–11.8 µm long, 4.5–6.1 µm wide, ratio length/width 1.3–2.6 (puzzle-shaped), 8.0– 9.9 µm long, 2.9–4.3 µm wide, ratio length/width 2.1–3.3 (rectangular); scales of inner side 11.8–19.4 µm long, 4.5–7.4 µm wide, ratio length/width 2.3–2.9 (gem-like), 15.1–17.7 µm long, 4.2–6.6 µm wide, ratio length/width 2.3–4.2 (leafshaped).

RADULA. Up to 37 overcrowded and undifferentiated teeth per row, comb-like and usually with five elongated cusps, but occasionally four or six/seven cusps, outermost cusps 39–77% of length of inner cusps; last marginal teeth with four to five triangular and pointed cusps, outer ones and median cusp (cusps 1, 3 and 5, when five cusps present) usually 53–74% of length of larger cusps; teeth with four cusps 4.4–4.7 µm wide, teeth with five cusps 3.7–5.5 µm wide (teeth in the center of row slightly wider than marginal teeth), teeth with six/seven cusps 6.0– 6.6 µm wide, last marginal teeth 3.0– 4.2 µm wide.


Inella  ” harryleei has a different radular morphology when compared to Inella obliqua  (May, 1915), which is the single species with illustrated soft parts of Inella  s.s. ( Marshall 1983). The operculum of “ I ”. harryleei is slightly eccentric ( Fig. 30CView Fig), similarly to I. obliqua  and S. gaesona  .

The undifferentiated and overcrowded teeth of “ I ”. harryleei resemble those of Monophorus olivaceus  ( Figs 17–18View FigView Fig) and M. erythrosoma  , with a typical tooth bearing five comb-like cusps, in spite of slight variations in the number of cusps. Just like M. olivaceus  , the last marginal teeth of “ I ”. harryleei are considerably reduced in size. Species of Monophorus  are usually sampled at mid-low depths of the continental shelf ( Fernandes & Pimenta 2017a), considering that “ I ”. harryleei similarly inhabits depths of 63–90 m ( Rolán & Fernández-Garcés 2008; present study). However, “ I ”. harryleei shows significant differences in the shell (e.g., abapical spiral cord of teleoconch much developed) and operculum (with an ovoid shape, almost elliptical, and nucleus more eccentric) compared to Monophorus  . Although it seems premature to reallocate “ I ”. harryleei in Monophorus  , “ I ”. harryleei is certainly related to the informal “ Inella  group” ( Marshall 1983).


Florida Museum of Natural History