Edessa sexdens, Fabricius, 1803

Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da & Fernandes, José Antônio Marin, 2023, Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa), Zootaxa 5372 (1), pp. 1-128 : 6-10

publication ID

https://doi.org/ 10.11646/zootaxa.5372.1.1

publication LSID

lsid:zoobank.org:pub:1300D562-3E0B-4F72-933F-8FA7D28F6853

DOI

https://doi.org/10.5281/zenodo.10574083

persistent identifier

https://treatment.plazi.org/id/414B87B5-FFD6-9843-FF63-D95A708FFAC9

treatment provided by

Plazi

scientific name

Edessa sexdens
status

 

This key includes all species of the E. sexdens group and was made based on the combination of body characteristics, should be noted that living specimens may have a slightly different color as noted above.

1 Body predominantly orange to red with large black areas on pronotum, coria and ventral surface of abdomen (see Mendonça et al., 2023: Fig. 27 E–F View FIGURE 27 )......................................................................... E. miniata

- Body predominantly green to brown with dark marks usually present only on humeral angles, connexivum and bands on ventral surface of abdomen ( Fig. 56 A–F View FIGURE 56 )........................................................................ 2

2 Ventral surface of the body with alternated yellow to greenish-yellow and green bands ( Figs. 56 B View FIGURE 56 , 59 B View FIGURE 59 , 65 D View FIGURE 65 , 70 B, F View FIGURE 70 ; see Mendonça et al., 2023: Fig. 9 B View FIGURE 9 , in poorly preserved specimens, greenish-yellow bands turn yellow and green bands turn brown).............................................................................................. 3

- Ventral surface of the body brown or yellow to orange, usually with dark bands, sometimes without bands ( Fig. 55 D, F View FIGURE 55 ).... ................................................................................................... 9

3 Humeral angles slightly curved and robust ( Figs. 56 A View FIGURE 56 , 59 A View FIGURE 59 , 67 E View FIGURE 67 ).............................................. 4

- Humeral angles curved and narrow ( Figs. 65 C View FIGURE 65 , 70 A, E View FIGURE 70 ; see Mendonça et al., 2023: Fig. 9 A View FIGURE 9 )........................ 6

4 All legs reddish brown; antennae reddish-brown ( Figs. 56 B View FIGURE 56 , 67 F View FIGURE 67 ).............................................. 5

- Hindlegs yellow to dark yellow, remaining legs reddish-brown; antennae black ( Fig. 59 A–B View FIGURE 59 )....... E. flavofemorata sp. n.

5 Second antennal segment shorter than third ( Fig. 67 E–F View FIGURE 67 )....................................... E. rufofusca sp. n.

- Second antennal segment longer than third ( Fig. 56 A–B View FIGURE 56 )................................... E. chloroicterica sp. n.

6 Scutellum with rounded apex ( Fig. 65 C View FIGURE 65 ; see Mendonça et al., 2023: Fig. 9 A View FIGURE 9 )..................................... 7

- Scutellum with acute apex ( Fig. 70 A, E View FIGURE 70 )................................................................... 8

7 Pronotum with anterolateral margin concave ( Fig. 65 C View FIGURE 65 ); arms of anterior bifurcation the metasternal process narrow, deep ( Fig. 35 G View FIGURE 35 )............................................................................ E. nigroantennata sp. n.

- Pronotum with anterolateral margin slightly concave; arms of anterior bifurcation the metasternal process with swollen, shallow (see Mendonça et al., 2023: Fig. 9 A–B View FIGURE 9 ).......................................................... E. imbecilla

8 Arms of anterior bifurcation of the metasternal process shallow ( Fig. 49 F View FIGURE 49 )........................ E. tuberculata sp. n.

- Arms of anterior bifurcation of the metasternal process deep ( Fig. 51 G View FIGURE 51 )......................... E. viridifasciata sp. n.

9 Humeral angles long, straight, somewhat frontward directed, posterior margin crenulate; posterior pronotal angles with a triangular, tooth-like projection ( Fig. 52 A, D–E View FIGURE 52 ; see Mendonça et al., 2023: Fig. 25 A View FIGURE 25 )............................. 10

- Humeral angles long or short, sinuous or almost straight, distally curved backward, posterior margin smooth; posterior pronotal angles without projections ( Figs. 55 C View FIGURE 55 , 58 A View FIGURE 58 ).............................................................. 11

10 Posterior pronotal tooth-like projections reduced; apex of humeral angles dark brown (see Mendonça et al., 2023: Fig. 25 A View FIGURE 25 )........................................................................................ E. cylindricornis

- Posterior pronotal tooth-like projections strongly developed; apex of humeral angles yellow ( Fig. 52 A, D–E View FIGURE 52 )............................................................................................... E. magnifica comb. n.

11 Connexival segments without dark spots ( Figs. 55 C View FIGURE 55 , 58 A View FIGURE 58 )................................................... 12

- Connexival segments with a pair of dark spots ( Figs. 62 E View FIGURE 62 , 64 A View FIGURE 64 , 66 E View FIGURE 66 ).......................................... 27

- Connexival segments almost entirely black ( Fig. 54 E View FIGURE 54 ).......................................... E. amaurata sp. n.

12 Evaporatorium dark, contrasting with surrounding surface ( Figs. 55 D View FIGURE 55 , 58 B View FIGURE 58 ; see Mendonça et al., 2023: Figs. 26 B, D View FIGURE 26 , 27 D View FIGURE 27 , 29 B View FIGURE 29 ).............................................................................................. 13

- Evaporatorium concolorous with surrounding surface ( Figs. 54 B View FIGURE 54 , 62 B View FIGURE 62 , 69 F View FIGURE 69 ).................................... 18

13 Humeral angles barely developed, at least 1.7 times wider than long ( Figs. 55 C View FIGURE 55 , 58 A View FIGURE 58 ; see Mendonça et al., 2023: Fig. 26 A–B View FIGURE 26 ).............................................................................................. 14

- Humeral angles developed, from 1.5 times wider than long to clearly longer than wide (see Mendonça et al., 2023: Figs. 26 C View FIGURE 26 , 27 C View FIGURE 27 , 29 A View FIGURE 29 )......................................................................................... 16

14 Pronotal disc coarse; ventral intersegmental bands of abdomen narrow, solid dark brown (see Mendonça et al., 2023: Fig. 26 A–B View FIGURE 26 )...................................................................................... E. gentilitia

- Pronotal disc smooth; ventral intersegmental bands of abdomen narrow or wide, diffused brown ( Figs. 55 C–D View FIGURE 55 , 58 A–B View FIGURE 58 ).... .................................................................................................. 15

15 Legs dark-brown; ventral intersegmental bands of abdomen very wide and dark brown ( Fig. 55 D View FIGURE 55 )..... E. brevicornis sp. n.

- Legs yellow to dark-yellow; ventral intersegmental bands of abdomen narrow and brown ( Fig. 58 B View FIGURE 58 )..... E. flavilinea sp. n.

16 Humeral angles wider than long (1.3 times wider than long), apex brown (see Mendonça et al., 2023: Fig. 29 A View FIGURE 29 ).................................................................................................. E. phoenicopus

- Humeral angles longer than wide, apex black or yellow (see Mendonça et al., 2023: Figs. 26 C View FIGURE 26 , 27 C View FIGURE 27 )................. 17

17 Antennae yellow; humeral angles pale yellow; corial veins yellow; ventral intersegmental bands of abdomen diffused brown (see Mendonça et al., 2023: Fig. 27 C–D View FIGURE 27 )............................................................ E. lewisi

- Antennae reddish brown; humeral angles black; corial veins green, paler than the surrounding surface; ventral intersegmental bands of abdomen solid black (see Mendonça et al., 2023: Fig. 26 C–D View FIGURE 26 )............................... E. helicoceras

18 Humeral angles brown ( Fig. 54 A View FIGURE 54 ; see Mendonça et al., 2023: Figs. 25 E View FIGURE 25 , 27 A View FIGURE 27 )................................... 19

- Humeral angles black ( Figs. 62 A View FIGURE 62 , 69 E View FIGURE 69 ).................................................................. 21

19 Ventral surface of the body with narrow black bands on thorax and abdomen (see Mendonça et al., 2023: Figs. 25 F View FIGURE 25 , 27 B View FIGURE 27 ).. .................................................................................................. 20

- Ventral surface of thorax and abdomen with wide brown bands ( Fig. 54 B View FIGURE 54 )............................ E. accola sp. n.

20 Humeral angles long (1.3 times wider than long); ventral abdomen without black spot close to each spiracle (see Mendonça et al., 2023: Fig. 25 E–F View FIGURE 25 ).......................................................................... E. fallenii

- Humeral angles short (1.6 times wider than long); ventral abdomen with black spot close to spiracles on each segment (see Mendonça et al., 2023: Fig. 27 A–B View FIGURE 27 )............................................................. E. leucocera

21 Arms of anterior bifurcation of metasternal process barely expanded laterally at apex ( Figs. 25 G View FIGURE 25 , 48 G View FIGURE 48 )............... 22

- Arms of anterior bifurcation of metasternal process expanded laterally at apex ( Figs. 10 G View FIGURE 10 , 40 G View FIGURE 40 )..................... 24

22 Humeral angles short and slightly curved backward ( Fig. 62 A View FIGURE 62 ; see Mendonça et al., 2023: Fig. 28 A View FIGURE 28 )................. 23

- Humeral angles long and curved backward ( Fig. 69 E View FIGURE 69 )..................................... E. truncatiacantha sp. n.

23 Femora darker than the rest of the leg (see Mendonça et al., 2023: Fig. 28 A–B View FIGURE 28 ).......................... E. nigriclava

- Femora concolorous with the rest of the leg ( Fig. 62 B View FIGURE 62 )....................................... E. immaculata sp. n.

24 Propleural band black and long, almost reaching base of humeral angles ( Figs. 67 B View FIGURE 67 , 70 D View FIGURE 70 ).......................... 25

- Propleural band brown and short, slightly passing propleural sulcus ( Figs. 57 B View FIGURE 57 , 69 B View FIGURE 69 )............................. 26

25 Ventrally prothorax with anterolateral margin gently grooved, humeral angles black spot broad, propleura shallowly punctured, area between pro and mesopleura brown ( Fig. 67 B View FIGURE 67 )............................................. E. redunca sp. n.

- Ventrally prothorax with anterolateral margin strongly grooved, humeral angles black spot narrow, propleura deeply punctured, area between pro and mesopleura black ( Fig. 70 D View FIGURE 70 )........................................ E. ventrocarinata sp. n.

26 Dorsal surface grass green ( Fig. 57 A View FIGURE 57 ); ventrally prothorax with humeral angles black spot narrow ( Fig. 57 B View FIGURE 57 ); arms of anterior bifurcation of metasternal process laterally expanded at apex ( Fig. 10 G View FIGURE 10 ); legs yellow ( Fig. 57 B View FIGURE 57 )....... E. decolorata sp. n.

- Dorsal surface olive green ( Fig. 69 A View FIGURE 69 ); ventrally prothorax with humeral angles black spot broad ( Fig. 69 B View FIGURE 69 ); arms of anterior bifurcation of metasternal process laterally well expanded at apex ( Fig. 46 G View FIGURE 46 ); legs reddish ( Fig. 69 B View FIGURE 69 ).................................................................................................... E. sphaerocornis sp. n.

27 Connexival dark spots not extended ventrally ( Figs. 63 B View FIGURE 63 , 64 B View FIGURE 64 , 66 D View FIGURE 66 ; see Mendonça et al., 2023: Figs. 19 B View FIGURE 19 , 26 F View FIGURE 26 , 28 D View FIGURE 28 )... .................................................................................................. 28

- Connexival dark spots extended ventrally ( Figs. 57 F View FIGURE 57 , 60 B View FIGURE 60 , 65 B View FIGURE 65 ).............................................. 37

28 Humeral angles as long as wide or wider than long ( Figs. 63 A View FIGURE 63 , 64 A View FIGURE 64 , 66 C View FIGURE 66 )...................................... 29

- Humeral angles longer than wide (see Mendonça et al., 2023: Fig. 26 E View FIGURE 26 ).................................... E. helix

29 Humeral angles wider than long ( Figs. 64 A View FIGURE 64 , 66 C View FIGURE 66 ; see Mendonça et al., 2023: Figs. 19 A View FIGURE 19 , 28 C View FIGURE 28 )..................... 30

- Humeral angles as long as wide ( Figs. 63 A View FIGURE 63 , 64 C View FIGURE 64 ).......................................................... 33

30 Humeral angles laterally directed, apex clearly globose, robust (see Mendonça et al., 2023: Fig. 28 C View FIGURE 28 )....... E. nigricornis

- Humeral angles posteriorly directed, small, apex slightly globose ( Figs. 64 A View FIGURE 64 , 66 C View FIGURE 66 ; see Mendonça et al., 2023: Fig. 19 A View FIGURE 19 )... .................................................................................................. 31

31 Black spots of connexivum small and narrow ( Fig. 66 C View FIGURE 66 ; see Mendonça et al., 2023: Fig. 19 A View FIGURE 19 )...................... 32

- Black spots of connexivum large and wide ( Fig. 64 A View FIGURE 64 )....................................... E. maranhensis sp. n.

32 Apex of scutellum reaching the apex of coria ( Fig. 66 C View FIGURE 66 ); ventrally abdomen with narrow bands on intersegmental areas and pseudosutures ( Fig. 66 D View FIGURE 66 ).................................................................. E. proxima sp. n.

- Apex of scutellum not reaching the apex of coria (see Mendonça et al., 2023: Fig. 19 A View FIGURE 19 ); ventrally abdomen with wide bands on intersegmental areas and pseudosutures (see Mendonça e t al., 2023: Fig. 19 B View FIGURE 19 )......................... E. schirmeri

33 Body dorsally olivaceous to dark-brown, ventrally orange; solid black thoracic ventral bands ( Fig. 63 A–B View FIGURE 63 ).................................................................................................... E. infuscata sp. n.

- Body dorsally green, ventrally greenish-yellow; diffused brown thoracic bands ( Figs. 55 A–B View FIGURE 55 , 59 C–D View FIGURE 59 , 61 C–D View FIGURE 61 , 64 C–D View FIGURE 64 )... .................................................................................................. 34

34 Body ventrally with bands wide and somewhat vivid ( Fig. 64 D View FIGURE 64 )............................... E. maurocornis sp. n.

- Body ventrally with bands narrow and faded ( Figs. 55 B View FIGURE 55 , 59 D View FIGURE 59 , 61 D View FIGURE 61 )........................................... 35

35 Arms of anterior bifurcation of metasternal process shallow, receiving only the fourth rostral segment ( Fig. 17 G View FIGURE 17 )............................................................................................ E. fuliginocornis sp. n.

- Arms of anterior bifurcation of metasternal process deep, receiving the fourth and part of the third rostral segments ( Figs. 4 G View FIGURE 4 , 23 B View FIGURE 23 ).............................................................................................. 36

36 Arms of anterior bifurcation of metasternal process barely expanded at apex ( Fig. 4 G View FIGURE 4 )................ E. atricornis sp. n.

- Arms of anterior bifurcation of metasternal process well expanded at apex ( Fig. 23 B View FIGURE 23 )............. E. illuminocornis sp. n.

37 Evaporatorium black ( Fig. 57 F View FIGURE 57 )............................................................. E. fabricii sp. n.

- Evaporatorium concolorous with the surrounding surface ( Figs. 56 D View FIGURE 56 , 68 D View FIGURE 68 ; see Mendonça et al., 2023: Fig. 29 B View FIGURE 29 )....... 38

38 Anterolateral margins of pronotum and anterior margin of cicatrices with row of black punctures covered by solid black line; apex of scutellum exceeding end of coria (see Mendonça et al., 2023: Fig. 29 C View FIGURE 29 )............................ E. sexdens

- Anterolateral margins of pronotum and anterior margin of cicatrices with row of concolorous to blackish punctures never covered by solid black line; apex of scutellum not reaching or level with end of coria ( Figs. 56 C View FIGURE 56 , 57 C View FIGURE 57 , 58 C View FIGURE 58 , 68 C View FIGURE 68 ; see Mendonça et al., 2023: Fig. 24 A, E View FIGURE 24 )..................................................................... 39

39 Humeral angles dorsally concolorous or lighter than pronotum ( Figs. 56 C View FIGURE 56 , 60 C View FIGURE 60 , 64 E View FIGURE 64 , 68 C View FIGURE 68 ; see Mendonça et al., 2023: Figs. 24 A, E View FIGURE 24 , 29 E View FIGURE 29 )....................................................................................... 40

- Humeral angles dorsally dark-brown to black, contrasting with pronotum ( Figs. 57 C View FIGURE 57 , 58 E View FIGURE 58 ; see Mendonça et al., 2023: Figs. 22 A View FIGURE 22 , 24 C View FIGURE 24 )......................................................................................... 46

40 Humeral angles straight, laterally directed ( Fig. 68 C View FIGURE 68 ; see Mendonça et al., 2023: Fig. 24 E View FIGURE 24 )......................... 41

- Humeral angles curved backward ( Figs. 56 C View FIGURE 56 , 60 C View FIGURE 60 , 64 E View FIGURE 64 ; see Mendonça et al., 2023: Figs. 24 A View FIGURE 24 , 29 E View FIGURE 29 )................ 42

41 Apex of humeral angles not curved; spots of connexival segments small ( Fig. 68 C View FIGURE 68 )................... E. simulata sp. n.

- Apex of humeral angles slightly curved; spots of connexival segments large (see Mendonça et al., 2023: Fig. 24 E View FIGURE 24 )................................................................................................... E. congrua

42 Humeral angles acuminated, small, not globose ( Fig. 56 C View FIGURE 56 ; see Mendonça et al., 2023: Fig. 29 E–F View FIGURE 29 ).................. 43

- Humeral angles rounded, somewhat developed, slightly globose ( Figs. 60 C View FIGURE 60 , 64 E View FIGURE 64 ; see Mendonça et al., 2023: Fig. 24 A View FIGURE 24 ).... .................................................................................................. 44

43 Ventral body surface, antennae and legs reddish yellow (see Mendonça et al., 2023: Fig. 29 F View FIGURE 29 ); arms of anterior bifurcation the metasternal process shallow (see Mendonça et al., 2023: Fig. 23 G View FIGURE 23 )........................................ E. urus

- Ventral body surface, antennae and legs yellowish ( Fig. 56 D View FIGURE 56 ); arms of anterior bifurcation the metasternal process deep ( Fig. 8 B View FIGURE 8 ).................................................................................. E. cornicula sp. n.

44 Dark spots of connexival segments large, occupying most of the segment ( Fig. 64 E View FIGURE 64 ; see Mendonça et al., 2023: Fig. 24 A View FIGURE 24 ); body ventral surface orange or dark yellow to brown ( Fig. 64 F View FIGURE 64 ; see Mendonça et al., 2023: Fig. 24 B View FIGURE 24 )................ 45

- Dark spots of connexival segments small, occupying only the lateral corners ( Fig. 60 C View FIGURE 60 ); body ventral surface yellow ( Fig. 60 D View FIGURE 60 )............................................................................... E. guatemalensis sp. n.

45 Humeral angles long (see Mendonça et al., 2023: Fig. 24 A View FIGURE 24 ); arms of anterior bifurcation the metasternal process divergent and shallow (see Mendonça et al., 2023: Fig. 1 G View FIGURE 1 ).......................................................... E. alces

- Humeral angles short ( Fig. 64 E View FIGURE 64 ); arms of anterior bifurcation the metasternal process slightly divergent and very shallow ( Fig. 33 F View FIGURE 33 )................................................................................... E. mimica sp. n.

46 Ventral surface of abdomen with bands of intersegmental areas solid black, not interrupted medially, except in segments II–III (see Mendonça et al., 2023: Fig. 22 B View FIGURE 22 )........................................................... E. subandina

- Ventral surface of abdomen with bands of intersegmental areas darkened or absent on raised median area ( Figs. 57 D View FIGURE 57 , 58 F View FIGURE 58 ).. .................................................................................................. 47

47 Humeral angles as long as wide, sinuous, anterolaterally directed with distal part curved backward ( Fig. 58 E View FIGURE 58 ; see Mendonça et al., 2023: Fig. 24 C View FIGURE 24 ).................................................................................. 48

- Humeral angles wider than long, laterally or posterolaterally directed with distal part straight or curved backward ( Figs. 57 C View FIGURE 57 , 60 E View FIGURE 60 , 61 A View FIGURE 61 , 65 A View FIGURE 65 , 66 E View FIGURE 66 , 67 C View FIGURE 67 , 68 E View FIGURE 68 ; see Mendonça et al., 2023: Fig. 25 C View FIGURE 25 )....................................... 49

48 Antennae dark brown; humeral angles with yellow to orange band adjacent to black distal part (see Mendonça et al., 2023: Fig. 24 C View FIGURE 24 ).......................................................................................... E. bella

- Antennae pale yellow to orange; humeral angles concolorous with pronotum except the black distal part ( Fig. 58 E View FIGURE 58 )......................................................................................... E. flavoantennata sp. n.

49 Humeral angles laterally directed ( Figs. 57 C View FIGURE 57 , 60 E View FIGURE 60 , 61 A View FIGURE 61 , 65 A View FIGURE 65 , 67 C View FIGURE 67 ).......................................... 50

- Humeral angles posterolaterally directed ( Figs. 61 E View FIGURE 61 , 63 C, E View FIGURE 63 , 66 E View FIGURE 66 , 68 E View FIGURE 68 ; see Mendonça et al., 2023: Fig. 25 C View FIGURE 25 )......... 54

50 Humeral angles short; ventral black spot of humeral angles restricted to the apical margin ( Fig. 65 A–B View FIGURE 65 )................................................................................................... E. nigroacantha sp. n.

- Humeral angles long; ventral brown spot of humeral angles covering most of the angles ( Figs. 57 C–D View FIGURE 57 , 60 E–F View FIGURE 60 , 61 A–B View FIGURE 61 , 67 C–D View FIGURE 67 ).............................................................................................. 51

51 Humeral angles with dorsal black spot ( Figs. 57 C View FIGURE 57 , 67 C View FIGURE 67 )..................................................... 52

- Humeral angles with dorsal dark brown spot ( Figs. 60 E View FIGURE 60 , 61 A View FIGURE 61 )................................................ 53

52 Apex of humeral angles curved; femora darker than tibiae and tarsi ( Fig. 57 C–D View FIGURE 57 ); distribution: southeastern Brazil ( Fig. 72 View FIGURE 72 )........................................................................................ E. echyra sp. n.

- Apex of humeral angles slightly curved; tibiae and tarsi darker than femora ( Fig. 67 C–D View FIGURE 67 ); distribution: Panama ( Fig. 72 View FIGURE 72 )........................................................................................... E. reuteri sp. n.

53 Apex of humeral angles straight and swollen ( Fig. 61 A View FIGURE 61 ); arms of anterior bifurcation the metasternal process well expanded at apex ( Fig. 22 F View FIGURE 22 ); body ventrally with uniformly wide black bands on intersegmental areas ( Fig. 61 B View FIGURE 61 ).................................................................................................... E. humerorotunda sp. n.

- Apex of humeral angles slightly curved and slightly swollen ( Fig. 60 E View FIGURE 60 ); arms of anterior bifurcation the metasternal process expanded at apex ( Fig. 21 F View FIGURE 21 ); body ventrally with black bands on intersegmental areas tapering to both ends ( Fig. 60 F View FIGURE 60 )...................................................................................... E. humeroglobosa sp. n.

54 Pseudosutures of the abdomen with solid dark brown to black band ( Figs. 66 F View FIGURE 66 , 68 F View FIGURE 68 ).............................. 55

- Pseudosutures of the abdomen concolorous with the surrounding surface or light-brown, band smudged ( Figs. 61 F View FIGURE 61 , 63 D, F View FIGURE 63 ; see Mendonça et al., 2023: Fig. 25 D View FIGURE 25 ).................................................................... 56

55 Humeral angles long and slightly curved backward ( Fig. 68 E View FIGURE 68 ); ventral abdomen yellow with black spot close to spiracle on each segment ( Fig. 68 F View FIGURE 68 ); distribution: Suriname ( Fig. 72 View FIGURE 72 ).................................... E. surinamensis sp. n.

- Humeral angles short and curved backward ( Fig. 66 E View FIGURE 66 ); ventral abdomen reddish brown without black spots close to spiracles ( Fig. 66 F View FIGURE 66 ); distribution: Costa Rica ( Fig. 74 View FIGURE 74 )........................................ E. pseudonigroangulata sp. n.

56 Ventral intersegmental bands of abdomen not reaching the lateral spots in segments II–VI ( Figs. 59 F View FIGURE 59 , 61 F View FIGURE 61 , 62 D View FIGURE 62 , 63 F View FIGURE 63 , 65 F View FIGURE 65 , 68 B View FIGURE 68 ; see Mendonça et al., 2023: Fig. 25 D View FIGURE 25 ).................................................................. 57

- Ventral intersegmental bands of abdomen reaching the lateral spots in segments II–VI ( Figs. 62 F View FIGURE 62 , 63 D View FIGURE 63 )............... 63

57 Ventral surface of abdomen yellow ( Fig. 62 D View FIGURE 62 ; see Mendonça et al., 2023: Fig. 25 D View FIGURE 25 ).............................. 58

- Ventral surface of abdomen reddish yellow to reddish brown ( Figs. 59 F View FIGURE 59 , 61 F View FIGURE 61 , 63 F View FIGURE 63 , 65 F View FIGURE 65 , 68 B View FIGURE 68 )...................... 59

58 Humeral angles long with dorsal spot covering 3/4 of the humeral angles; ventral abdominal spine in segment III distally rounded ( Fig. 62 C–D View FIGURE 62 )....................................................................... E. inca sp. n.

- Humeral angles short with dorsal spot covering 2/3 of the humeral angles; ventral abdominal spine in segment III distally acuminated (see Mendonça et al., 2023: Fig. 25 C–D View FIGURE 25 ).................................................. E. electa

59 Dorsal surface green; humeral angles with dorsal black spot restricted to the angles; connexival spots yellow ( Figs. 61 E View FIGURE 61 , 63 E View FIGURE 63 )................................................................................................ 60

- Dorsal surface reddish brown; humeral angles with dorsal black spot extended to the disc of pronotum; connexival spots orange ( Figs. 59 E View FIGURE 59 , 65 E View FIGURE 65 , 68 A View FIGURE 68 )............................................................................... 61

60 Arms of anterior bifurcation of metasternal process well expanded at apex ( Fig. 24 B View FIGURE 24 ); ventral intersegmental bands of abdomen black ( Fig. 61 F View FIGURE 61 ).......................................................................... E. imitans sp. n.

- Arms of anterior bifurcation of metasternal process expanded at apex ( Fig. 30 G View FIGURE 30 ); ventral intersegmental bands of abdomen brown ( Fig. 63 F View FIGURE 63 )....................................................................... E. maculosa sp. n.

61 Ventral black spot of humeral angles covering most of the angles ( Fig. 59 F View FIGURE 59 ).................... E. fulvomaculata sp. n.

- Ventral black spot of humeral angles restricted to the margin of the angles ( Figs. 65 F View FIGURE 65 , 68 B View FIGURE 68 )......................... 62

62 Metasternal process with arms of anterior bifurcation receiving the fourth and part of the third rostral segment; arms of anterior bifurcation the metasternal process barely expanded laterally at apex ( Fig. 43 F View FIGURE 43 ); ventral abdominal spine in segment III acuminated ( Fig. 68 B View FIGURE 68 )..................................................................... E. rutilata sp. n.

- Metasternal process with arms of anterior bifurcation not completely receiving the fourth rostral segment; arms of anterior bifurcation the metasternal process expanded laterally at apex ( Fig. 36 G View FIGURE 36 ); ventral abdominal spine in segment III rounded ( Fig. 65 F View FIGURE 65 )............................................................................... E. nigrocrocata sp. n.

63 Lateral portion of each abdominal segment with the middle yellow spot smaller than the anterior and posterior black spots combined; ( Fig. 63 C–D View FIGURE 63 )................................................................... E. limeirai sp. n.

- Lateral portion of each abdominal segment with the middle yellow spot larger than the anterior and posterior black spots combined ( Figs. 54 C–D View FIGURE 54 , 55 E–F View FIGURE 55 , 56 E–F View FIGURE 56 , 58 C–D View FIGURE 58 , 60 A–B View FIGURE 60 , 62 E–F View FIGURE 62 ; see Mendonça et al., 2023: Figs. 10 F–G View FIGURE 10 , 28 E–F View FIGURE 28 )... 64

64 Dorsal black spot of humeral angles clearly expanded over the disc of pronotum; propleura with dark band reaching the base of humeral angles ( Figs. 54 C–D View FIGURE 54 , 58 C–D View FIGURE 58 ).................................................................. 65

- Dorsal black spot of humeral angles restricted to the angles; propleura with dark band not reaching the base of humeral angles ( Figs. 55 E–F View FIGURE 55 , 56 E–F View FIGURE 56 , 60 A–B View FIGURE 60 , 62 E–F View FIGURE 62 , 66 A–B View FIGURE 66 , 69 C–D View FIGURE 69 ; see Mendonça et al., 2023: Figs. 10 F–G View FIGURE 10 , 28 E–F View FIGURE 28 )........... 66

65 Humeral angles long, mostly black ventrally; propleura with black band ( Fig. 54 C–D View FIGURE 54 ).............. E. amapaensis sp. n.

- Humeral angles short, mostly brown ventrally; propleura with brown band ( Fig. 58 C–D View FIGURE 58 )............. E. flaviventris sp. n.

66 Ventral surface orange; connexivum with orange spots (see Mendonça et al., 2023: Fig. 10 F–G View FIGURE 10 )................ E. lavata

- Ventral surface yellow to dark yellow; connexivum with yellow spots ( Figs. 55 E–F View FIGURE 55 , 56 E–F View FIGURE 56 , 60 A–B View FIGURE 60 , 66 A–B View FIGURE 66 ; see Mendonça et al., 2023: Fig. 28 E–F View FIGURE 28 ).............................................................................. 67

67 Propleura coarse, deeply punctured ( Fig. 56 F View FIGURE 56 ; see Mendonça et al., 2023: Fig. 28 F View FIGURE 28 )............................... 68

- Propleura smooth, shallowly punctured ( Figs. 55 F View FIGURE 55 , 60 B View FIGURE 60 , 62 F View FIGURE 62 , 66 B View FIGURE 66 , 69 D View FIGURE 69 )...................................... 69

68 Humeral angles incrassate, ventral black spot restricted to the apical margin ( Fig. 56 E–F View FIGURE 56 )........... E. crassicornis sp. n.

- Humeral angles slender, ventral black spot covering the apex (see Mendonça et al., 2023: Fig. 28 E–F View FIGURE 28 )..... E. nigroangulata

69 Humeral angles with ventral black spot restricted to the apical margin; black spots of connexivum small ( Figs. 62 E–F View FIGURE 62 , 66 A–B View FIGURE 66 ).............................................................................................. 70

- Humeral angles with ventral black spot covering half or more of the angles; black spots of connexivum large ( Figs. 55 E–F View FIGURE 55 , 60 A–B View FIGURE 60 , 69 C–D View FIGURE 69 )....................................................................................... 71

70 Pronotum with shallow punctures; humeral angles long; ventral intersegmental bands of abdomen black ( Fig. 62 E–F View FIGURE 62 )........................................................................................... E. indiscreta sp. n.

- Pronotum with deep punctures; humeral angles short; ventral intersegmental bands of abdomen brown ( Fig. 66 A–B View FIGURE 66 )............................................................................................ E. omopalla sp. n.

71 Dorsal black spot of humeral angles reaching the baseline of the angles, ventral black spot covering more than half of the angles ( Fig. 60 A–B View FIGURE 60 ).......................................................................... E. glomerosa sp. n.

- Dorsal black spot of humeral angles not reaching the baseline of the angles, ventral black spot covering less than half of the angles ( Figs. 55 E–F View FIGURE 55 , 69 C–D View FIGURE 69 ).......................................................................... 72

72 Humeral angles swollen ( Fig. 69 C–D View FIGURE 69 ); arms of anterior bifurcation of metasternal process expanded at apex ( Fig. 47 F View FIGURE 47 )........................................................................................ E. tenuivittata sp. n.

- Humeral angles barely swollen ( Fig. 55 E–F View FIGURE 55 ); arms of anterior bifurcation of metasternal process barely expanded at apex ( Fig. 6 G View FIGURE 6 ).............................................................................. E. brevihumeralis sp. n.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Pentatomidae

Genus

Edessa

GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF