Uracanthus triangularis Hope, 1833

Thongphak, Duangrat & Wang, Qiao, 2007, Taxonomic revision of the longicorn beetle genus Uracanthus Hope 1833 (Coleoptera: Cerambycidae: Cerambycinae: Uracanthini) from Australia., Zootaxa 1569 (1569), pp. 1-139 : 18-20

Uracanthus triangularis Hope, 1833

(Figs 24, 62, 95)

Uracanthus triangularis Hope, 1833: 64 .— Hope, 1834: 108; Lacordaire, 1869: 391; Best, 1882: 35; Tepper, 1887: 38; Froggatt, 1893: 29, 1902: 710, 1923: 112; Gahan, 1893: 182; French, 1900: 135; Gurney, 1911: 58; Aurivillius, 1912: 147; Gallard, 1916: 113; Tillyard, 1926: 223; McKeown, 1947: 66; Duffy, 1963:112.

Stenochorus angustatus Boisduval, 1835: 475 .— Gahan, 1893: 182 (synonymy).

Mallocera angustata .– Laporte, 1840: 425.

Description

Male. Body length, 21.68–33.13 mm; width, 4.24–6.47 mm.

Colour (Fig. 24). Body reddish brown to blackish brown with head and thorax darker. Head with dense white or yellowish pubescence except median frontal groove and a longitudinal glabrous area on vertex. Pronotum with 2 longitudinal stripes of white pubescence on each side: 1 on disc, vague, and 1 near ventral side, distinct. Each elytron with a large clearly defined subtriangular glabrous mark, extending from shoulder to basal 1/3, clearly margined with dense white pubescence; apical area glabrous, margined with a distinct line of dense white pubescence; remaining part of elytral disc with dense and fairly long pubescence. Body colour varies from reddish brown to blackish brown. The glabrous mark on the elytra may be reduced or expanded from the basic pattern.

Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; distance between lower lobes of eyes 1.5–1.85 × as long as distance between upper lobes of eyes; distance between upper lobes of eyes 1.0–1.25 × as long as distance between eyes on ventral side; genal length 0.35–0. 44 × as long as head width immediately below eyes. Antennae about as long as body; segments 5–10 flattened and produced on one side at apex; apical ¼ of segment 11 slightly thinner than basal ¾.

Thorax and abdomen. Pronotum 1.14–1.46 × as long as width, rounded at side; posterior margin 1.07– 1.36 × as wide as anterior margin; pronotal disc binodulose in middle and rugose transversely throughout. Scutellum semicircular, with sparse pubescence. Elytra 4.1–5.07 × as long as prothorax and 3.7–4.15 × as long as shoulder width; each elytron with 3 very feeble carinae; basal half of elytra with sparse coarse punctures; apex widely emarginate and bispinose. Abdomen slender; apex of terminal sternite truncate or slightly rounded.

Male terminalia. Apex of ventral median lobe narrowly emarginate, and apex of dorsal lobe rounded; ventral lobe slightly longer than or as long as dorsal lobe. spined region of internal sac about 2.1–2.5 × as long as basal unspined regions; spined region divided into 2 sections with an unspined gap between sections; first section longer than unspined gap with dense long simple spines; second section with sparse short simple spines ( Fig. 62a). Eighth sternite obliquely truncate at sides, shallowly emarginate at apex, with long and fairly long setae arising terminally; ventral surface with cloud-like processes ( Fig. 62b). Eighth tergite truncate or shallowly emarginate at apex, with fairly dense short simple spines and multi-branched spines on dorsal surface ( Fig. 62c). Paramere 1.5–1.67 × as long as wide, cylindrical in shape, apex rounded with long and short setae ( Fig. 62d).

Female. Body length, 17.17–34.19 mm; width, 3.05–6.97 mm.

Similar to males, but differs in having antennae and legs distinctly shorter, more robust abdomen; elytra slightly longer and wider, 3.75–4.4 × as long as shoulder width and 4.09–5.13 × as long as prothorax. Apex of terminal sternite rounded.

Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs ( Fig. 62e). Spermatheca clearly curved; spermathecal gland long and arising at base ( Fig. 62f).

Distribution Northern, eastern, and southwestern coasts of Australia ( Fig. 95).

Biology

The following were recorded as the host plants of this species: Acacia decurrens , A. mearnsii [= A. mollissima ], A. penninervis , A. pycnantha , A. sophorae [= A. longifolia var. sophorae ], Acacia spp. , Boronia pinnata , Eriostemon australasius [= E. lanceolatus ], Banksia ericifolia ; B. integrifolia , Banksia spp. , Hakea gibbosa , H. nodosa , H. serice and Lomatia longifolia . Adults were found on Leptospermum flavescens and Angophora sp. flowers and Kunzea ambigua and Xanthorrhoea sp. foliage. Eggs are deposited in small branches of the host trees. Adults emerge from September to December, although eclosion occurs as early as June. They were collected from October to April. Adults are attracted to MV light.

Comments

This species closely resembles U. pallens but differs in having the pronotal disc rugose transversely throughout; elytral apex more widely emarginate and more strongly and acutely spinose, and the subtriangular marks on elytra broader.

Material examined

Holotype. ♀. New Holland (= Australia); no data; type coll. 1767; bearing a red type label with Hope’s hand writing ( HMO).

Other material examined. 114 ♂, 118 ♀. QLD: 1 ♂, 1 ♀, Stanthorpe (28°39'S, 151°56'E), i.1961, C. Watts ( SAM) ; 1 ♂, same locality but Sivian ( AM) ; 1 ♂, same locality but Von Weildt ( QM) ; 1 ♀, same locality but Dr. K. K. Spence ( AM) ; 2 ♀, same locality but 30.iii.1963, P. Kerridge ( UQIC) ; 1 ♀, same locality but 2.xi.1927, S. M. Watson ( QDPI) ; 3 ♀, same locality but xii.1964, E. Sutton Coll. ( QM) ; 1 ♀, as above but 2.iv.1926 ( QM) ; 1 ♂, 70 km SW of Greenvale (19°00'S, 144°58'E), 16–28.i.1995, A. J. Watts ( SAM) ; 1 ♀, Bowarrady Creek (25°07'S, 153°08'E), Fraser Island , 5.x.1989, L. D. Buddle ( AM) ; 2 ♀, Kuranda (16°49'S, 145°38'E), F. P. Dodd ( SAM) ; 2 ♂, Moreton Island (27°11'S, 153°24'E), Ben Ewa Campground, 22.iii.1998, at light, J. & A. Skevington ( UQIC) ; 1 ♂, Oranmore Range , 5.iii.1958, A. R. Briay ( QDPI) ; 1 ♂, Archer River Xring (13°25'S, 142°56'E), 11.v.1989, MV lamp, G. & A. Daniels ( UQIC) ; 1 ♀, 320 m of Isla Gorge National Park (25°11'S, 149°58'E), 3.x.1992, MV lamp, G. Daniels ( UQIC) ; 1 ♂, 26 km W of Fairview (15°35'S, 144°04'E), 24.v.1989, MV lamp, G. & A. Daniels ( UQIC) ; 1 ♂, 1 ♀, 10 km N of Caboolture (27°04'S, 152°58'E), Elimbah, 11.iv.1984, C. E. Hagan ( UQIC) ; 1 ♀, Eclipse Island , Coll. no. 39-1785 ( WAM) ; 1 ♂, QLD, 1.viii.1964, A. W. Asjary ( UQIC) ; 1 ♂, Wyberba , 5–7.xi.1969, at light, E. C. Dahms ( QM) ; 3 ♂, 1 ♀, Cooloola Forestry Area , camp milo, iv.1978, at light, I. D. Galloway & E. Gympie ( QDPI) ; 1 ♂, 1 ♀, same locality but Banksia dom, 3.iii.1970, open forest to light, E. Dahms ( QM) ; 2 ♀, Glen Aplin , SQ,1946, S. R. E. Brock Collection ( ANIC) ; 3 ♂, same locality as above but 1957, A. Gemmell, S. R. E. Brock Collection ( ANIC) ; 2 ♀, same locality as above but 1912–1954, A. Gemmell, G. G. L. Gooding Collection ( ANIC) ; 2 ♂, 2 km N by NW Jawalbinna (15º45'S, 144º15'E), 17.i.1994, at light, P. Zborowski & E. D. Edwards ( ANIC) ; 1 ♂, 4 km W by S of Cooktown , 21.v.1977, I. F. B. Common & E. D. Edwards ( ANIC) ; 1 ♂, 24 km E by N of Ravenshoe, Palmerston Nation Park (17º35'S, 145º43'E), 17.xi.1981, J. Balderson ( ANIC) ; 1 ♂, South Dunmore SF (27º38'S, 150º58'E), 26.xii.1998, at MV light, S. G. Watkins Collection ( ANIC) ; 1 ♂, 13 km N by NW of Longreach (23º135'S, 144º04'E), Darr River , 7.iv.1976, J. F. Feehan ( ANIC) ; 1 ♀, Toowoomba , i.1980, C. Brook ( NMV) . NT: 2 ♂, Daly Waters (16°16'S, 133°22'E), 20.i.1971, T. Weir & A. Allwood ( NTM) ; 1 ♂, October Creek (16°28'S, 135°09'E), 100 m E of Dally Water , 21.iii.1972, on Borroloola Rd, N. W. Forester ( NTM) ; 1♀, NT, Cattle Creek (16º32' S, 136º10'E), 54 km S byW of Borroloola, 27.x.1975, M. S. Upton ( ANIC) . NSW: 1 ♂, Engadine (34º 03' S, 151º 01' E), 10.i.1974, R. H. Mulder Collection ( AM) ; 1 ♂, as above but 14.xii.1974 ( AM) ; 1 ♂, as above but 17.xii.1958, H. Hughes ( AM) ; 1 ♀, as above but 10.i.1958 ( AM) ; 1 ♀, as above but 14.xii.1974 ( AM) ; 1 ♀, as above but 28.xi.1968 ( AM) ; 1 ♀, as above but 14.xii.1974 ( AM) ; 1 ♀, as above but 10.i.1970 ( AM) ; 1 ♀, as above but 6.iii.1955 ( AM) ; 1 ♀, as above but 12.x.1960, ( AM) ; 2 ♂, 1♀, 5mi, S. Mendooran (31º 47'S, 149º 17'E), 19.ii.1972, MV lamp, G. Daniels ( AM) ; 2 ♀, same locality but 17.ii.1972 ( AM) ; 2 ♂, 1 ♀, Helensburgh (34º11'S, 150º58'E), xi.1961, H. Osborne ( ASCU) ; 1 ♀, same data but xii.1961 ( ASCU) ; 1 ♀, same data but xii.1962 ( ASCU) ; 1 ♂, same data but xii.1963 ( ASCU) ; 1 ♂, West Pymble (33º45'S, 151º07'E), near Sydney, 6.iii.1986, D. J. Scambler ( AM) ; 1 ♂, Goonoo State Forest (32º04' S, 148º54'E), 5 mi S. Mendooran , 24.iv.1974, D. K. McAlpine ( AM) ; 1 ♂, same data but 23.iv.1974 ( AM) ; 1 ♀, Budthingeroo Ceek (33º53'S, 150º00'E), Kanangra Boyd National Park , 24.xii.1977, G. Daniels ( AM) ; 1 ♀, Kuringgai Chase National Park (33º36'S, 151º12'E), 16.x.1971, MV lamp, G. Daniels ( AM) ; 1 ♀, same locality, N. C. Llayd ( ASCU) ; 1 ♂, same locality but 5.iv.1973, A. B. Rose ( MAM) ; 2 ♀, Newcastle (33º55'S, 151º00'E), i.1988, Coll. no. K36095 ( AM) ; 1 ♀, Round Hill Reserve (33º00'S, 146º10'E), 23–29.iv.1968, R. Lossin ( AM) ; 2 ♂, 2 ♀, Sydney (33º51'S, 151º12'E) ( MAM) ; 1 ♂, 1 ♀, same locality but ( SAM) ; 1 ♂, same locality but xii.1938 (no abdomen) ( MAM) ; 1 ♀, same locality ( SAM) ; 1 ♂, same locality but W. du Boulay, E. F. du Boulay Collection ( SAM) ; 1 ♂, same locality but 9.iii.1907, W. B. G ( ASCU) ; 1 ♂, same locality but Dean ( UQIC) ; 1 ♂, same locality but Coll. no. K36095 ( AM) ; 1 ♀, same locality but xi.1911, C. Gibbons ( AM) ; 1 ♂, 1 ♀, Blue Mountains National Park (33º36'S, 150º25'E), 15.xii.1984, on Leptospermum, G. R. Brown (ASCU) ; 1 ♀, Blue Mountains National Park (33º44'S, 150º30'E), 1.xii.1947 ( SAM) ; 1 ♀, Billy Blue Mountain Nation Park (34º02'S, 149º27'E), 30.i.1980, N. W. Rodd ( AM) ; 1 ♀, Mt. Tomah (33º32' S, 150º25'E), 22.xi.1980, N. W. Rodd ( AM) ; 1 ♂, 1 ♀, Mt. Wilson (33º59'S, 149º21'E), 27.xi.1980, D. A. Doolan, D. A. Doolan Collection ( AM) ; 1 ♂, same locality but 6.xi.1923, Carno rear, A. Musgrave & T. Carepbull ( AM) ; 1 ♂, National Park , 16.x.1921, A. Musgrave, Coll. no. K 44380 ( AM) ; 1 ♀, 1.5 mile E. of Freshwater River, Iluka district , 17.i.1971, D. K. McAlpine & A. Hughes ( AM) ; 1 ♀, Mosman (33º13'S, 150º50'E),1923, E. Balton, Coll. no. K47516 ( AM) ; 1 ♂, Lane Cove (33°48'S, 151°10'E), 12.xii.1943, N. W. Rodd ( AM) ; 1 ♂, same as above but 7.iv.1973, G. R. Brown ( ASCU) ; 1 ♂, Dorrigo (30°19'S, 152°43'E), 7.x.1973, D. A. Doolan, D. A. Doolan Collection ( AM) ; 1 ♀, North Ryde (33°47'S, 151°07'E), 17.i.1967, D. A. Doolan, D. A. Doolan Collection ( AM) ; 1 ♂, Bobbin Head , 23.iv.1969, K. Ayers ( MAM) ; 1 ♂, Wahroonga , 8.xii.1969, A. B. Rose ( MAM) ; 1 ♂, Oranmore, Hokas , 9.xii.1968, S. Barker ( SAM) ; 1 ♀, near Rivertree (28°37'S, 152°18'E), 15– 19.xii.1969, E. Dahms ( QM) ; 1 ♂, Bardwell Park (33°56'S, 151°08'E), 21.i.1943, C. E. Chadwick ( ASCU) ; 1 ♀, Wahroonga , 26.xi.1952, R. Dobson ( NMV) ; 1 ♂, same data as above ( NMV) ; 1 ♀, Sublime land, 15.xi.1902 ( ASCU) ; 1 ♂, Hornsby , 3.iii.1978, A. Beattic ( ASCU) ; 1 ♀, Wilton , 12.i.1966, V. J. Robinson ( ASCU) ; 1 ♂, as above ( ASCU) ; 1 ♂, Good Friday Mountain , 7.i.1968, on Fire trail, V. J. Robinson ( ASCU) ; 1 ♂, London Falls , 12.xii.1948, on Leptospermum flavescens flowers, C. E. Chadwick ( ASCU) ; 1 ♀, Botany , 13.xi.1994, Damagein ( ASCU) ; 2 ♂, 1 ♀, Gibraltar Range (29°33'S, 152°17'E), 3,000 ft. via Glen Innes , 27– 29.xii.1972, open forest, G. B. Monteith ( UQIC) ; 1 ♀, Bundjalung, National Park near Iluka , 12.iv.1981, M. J. Fletcher & G. R. Brown ( ASCU) ; 1 ♂, Canberra , 29.ii.1967, light trap, BCP ( WINC) ; 1 ♀, 8 km S of Mendooran (31°50'S, 149°06'E), 28.i.1987, G. & A. Daniels ( UQIC) ; 1 ♂, NSW ( SAM) ; 1 ♂, as above ( WADA) ; 1 ♂, same locality but I. D by lea ( WADA) ; 1 ♀, same locality ( MAM) ; 1 ♀, same locality ( MAM) ; 1 ♀, same locality but 26.xi.1952, Nat Mus, R. Dobson ( NMV) ; 1 ♀, same locality but 15.xii.1977, Rear Beach, A. D. Austin ( WINC) ; 1 ♀, Hornby , xi.1910, G. Gibbons ( AM) ; 1 ♀, Vaucluse , 19.ii.1920, J.M., Coll. no. K63681 ( MAM) ; 1 ♂, North Sydney , 11.viii.1946, P. Hadlington ( FCNI) ; 1 ♂, Lucas Heights, Sydney , NSW, 23.xi.1979, on Angophora bloom, D. C. Carne ( ANIC) ; 1 ♀, Sydney , NSW, R. W. Ferguson Collection ( ANIC) ; 1 ♂, Ebenezer, Sydney , 5.iii.1987, MV light, J. C. Keast ( FCNI) ; 1 ♂, Bonnet Bay, Sydney , 28.ii.1987, at light, L. S. William ( FCNI) ; 1 ♀, Bargo , 8.xii.1987, adult on Kunzea ambigua foliage, G. A. Webb ( FCNI) ; 1 ♂, Tubbamurra Creek, Barrington Tops , 4100 ft, 11.i.1956, K. D. Fairrey ( FCNI) ; 1 ♂, Heathcote , 10.xii.1979, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♀, same data as above but 2.xii.1978 ( ANIC) ; 1 ♂, same data as above but 4.xii.1977 ( ANIC) ; 2 ♂, 3 km N of Mt. Victoria ,, 22.xii.1978, on Xanthorrhoea, S. G. Watkins Collection ( ANIC) ; 1 ♂, Mt. York , NSW, 30.xi.1977, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♀, same data as above but 28.xi.1979 ( ANIC) ; 1 ♂, 50 miles S of Singleton , 5.i.1956, I. F. B. Common ( ANIC) ; 2 ♀, 20 miles NW of Upper Colo , 8.xi.1955, T. G. Campbell ( ANIC) ; 1 ♀, Mt Boyce , 16.xii.1972, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♂, 9 km N by NE of Coonabarabran , 24.x.1980, E. Britton ( ANIC) ; 1 ♀, Berkshire Park , 19.xi.1972, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♀, Hartley Vale , 27.xi.1977, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♀, Waterfall , NSW, 26.xii.1974, S. Watkins, S. G. Watkins Collection ( ANIC) ; 1 ♀, Kiata , 8.xii.1964, N. Dobrotworsky ( ANIC) ; 1 ♀, Wahroonga, H. J. Carter ( ANIC) ; 1 ♀, Toganmain , 4.x.1966, I. Rawley ( ANIC) ; 1 ♂, 1♀, National Park , xi.1952, J. G. Brook ( ANIC) ; 1 ♀, NSW, H. E. Cox Collection, present 1916 by Mrs Cox ( HMO) ; 1 ♀, Richmond R., New Holland, Miers Collection, presented 1880 by W. Miers ( HMO) ; 1 ♂, Glebe Park , 11.iv.1936, E. Sutton ( QM) ; 1 ♀, Mittagong , 20.iv.1922, H. J. Carter Coll. ( NMV) ; 1 ♂, Deewhy , ii.1928 ( ANIC) . ACT: 2 ♂, 3 ♀, Black Mountain , 14.ii.1966, at light trap, P. B. Carne ( ANIC) ; 2 ♂, 2 ♀, same locality as above but xi.1965 – ii.1966, E. B. Britton ( ANIC) ; 1 ♂, same locality as above but 28.ii.1969, at light trap ( ANIC) ; 1 ♂, same locality as above but 25–26.xii.1964, I. F. B. Common ( ANIC) ; 1 ♂, same locality as above but 16.ii.1968, M. S. Upton ( ANIC) ; 1 ♂, same locality as above but 15.ii.1936, A. D. Butcher ( NMV) ; 1 ♀, same data as above but 27.i.1936 ( NMV) ; 2 ♀, Lees Creek near Picadilly circus, 8.ii.1984, Calder & Stevens ( ANIC) . VIC: 1 ♂, 3 km S. Murchison , 8.i.1979, N. W. Rodd ( AM) ; 1 ♂: Mt. Difficult (37°00'S, 142°26'E), Grampians, 2600 ft., 7.ii.1956, N. B. Tindale ( SAM) ; 1 ♀, Melbourne , 9.xi.1932, F. E. Wilson Collection ( NMV) ; 2 ♂, 10 miles N of Wingan Inlet , 3.xi.1969, I. F. B. Common ( ANIC) ; 1 ♀, 17 miles NW Orbost , 8.xii.1956, E. F. Riek ( ANIC) ; 1 ♂, Beechworth, C. Oke ( NMV) ; 1 ♀, Abbeyard , 27.ix.1960 ( NMV) ; 1 ♂, Mordialloc (no abdomen) ( NMV) ; 1 ♂, Victoria ( WADA) ; 1 ♂, Victoria ( AM) . WA: 1 ♀, Denmark (34°58'S, 117°21'E), i.1997, M. & L. Jone ( WAM) ; 1 ♀, same data but i.1998 ( WAM) ; 1 ♂, same locality but 24.xi.1999, R. P. McMillan ( WAM) ; 1 ♂, Haker sterm, Turle Rock, Muckenburra , 16.x.1979, R. P. McMillan ( WAM) ; 1 ♂, Esperance (33°51'S, 121°53'E), 26.ix.1970, A. V. Thomas ( WAM) ; 1 ♀, Wurarga (28°24'S, 116°17'E), Coll. no. 37-4439 ( WAM) ; 1 ♀, Karridale (34°12'S, 115°06'E), 18.xii.1964, L. M. Olkalloran ( QDPI) ; 1 ♀, WA ( WADA) . Locality Unknown: 1 ♂, E. C. Vallis Bequest, Don. vii.1965 ( QM) ; 1 ♂, National Park , T. Mec. 13.x.1911 ( ASCU) ; 1 ♂, Nation Pk. , T. Mec. 13.x.1911 ( ASCU) ; 1 ♂, R. R. McMillan Collection ( WAM) ; 1 ♂, Coll. no. 2926 ( SAM) ; 1 ♂, 10.ii.1920, Coll. no. K63681 ( AM) ; 1 ♀, no data ( NMV) ; 1 ♀, Relton Bequest , Coll. no. 3455 ( QM) ; 1 ♀, no data ( ASCTO) ; 1 ♀, no data ( WADA) ; 1 ♀, 11.x.1915, bred from Lomatia longifolia, J. C. Dixon (NMV) ; 1 ♀, A. H. Elston collection ( AM) ; 1 ♀, 29.iii.1967, D. A. Doolan, D. A. Doolan Collection ( AM) ; 1 ♀, N. RPC, 17.iv.1967, D. A. Doolan Collection ( AM) ; 1 ♂, 1 ♀, K. G. Sound ( MAM) ; 1?, no data (no abdomen) (SAM); 1 ♂, Deewhy , ii.1928 ( ANIC) ; 1 ♀,? Park Coll , 1924, WWF ( ANIC) ; 1 ♀, no locality label, S. R. E. Brock Collection ( ANIC) ; 1 ♀, ix.1907, ex Acacia mollissima (NMV) ; 1 ♂, Slud Park (?), Acacia mollissima , x.1907 ( NMV) ; 1 ♀, i.1908, Acacia (NMV) ; 2 ♀, ix.1913, Acacia (NMV) ; 1 ♂, xii., Acacia (NMV) ; 1 ♀, out of black wattle ( Acacia mearnsii ), S. Park ( WINC) ; 1 ♀, same as above but ( NMV) ; 1 ♂, 2 ♀, no label ( NMV) .