Ayacucho spielbergi, Benedetti & Pinto-da-Rocha, 2022

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

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Ayacucho spielbergi

sp. nov.

3.20. Ayacucho spielbergi sp. nov.

Figs 3A View Figure 3 , 7O, P View Figure 7 , 12G, H View Figure 12 , 22H View Figure 22 , 28 View Figure 28


MALE: Measurements (n =3) DSW: 4.0-4.9 (4.9); DSL: 4.6-5.0 (5.0); CL: 1.6-2.0 (2.0). FIVL: 7.6-8.6 (8.6). ChL: 2.1-3.9 (3.9). Coloration: (Fig. 22H View Figure 22 ) Carapace, lateral margins of DS, spines of scutal area III and legs dark brown. Reddish chelicera. Pedipalpus yellowish with black spots. Posterior margin of DS and free tergites I-III light yellow. Scutal areas light yellow, with series of dark spots. Dorsum: (Fig. 3A View Figure 3 ) Gamma-type DSS. Anterior margin of the DS with a median elevation, with few granules sparsely distributed. Ocularium with medial depression; with a pair of spines and few granules near the eyes. Areas I-IV with few granules concentrated in the lateral regions; area III with a pair of spines directed backwards, located at the highest point in elevations of the integument, totally granular; area IV with a pair of small lateral tubercles. Posterior margins of DS smooth, with a pair of medium tubercles. Free tergites I-II with a pair of median tubercles; and a lateral pair of tiny tubercles. Free tergite III with two elongated median tubercles, larger than those of the free tergites I-II. Chelicerae: (Fig. 3A View Figure 3 ) Swollen in large males (swollen in holotype); slightly larger than that of the females and small males. Segment I granulate; II with few granules; III with two teeth. Pedipalpus : Trochanter with two ventroapical setiferous tubercles, one large and one smaller. Femur with a row of six ventral setiferous tubercles in the basal ⅔. Patella with a small retroapical tubercle. Tibia: prolateral iiIii, retrolateral IIi. Tarsus: prolateral iiIiIi, retrolateral iIiIi. Venter: Coxa I with a row of 4-5 tubercles. Coxae II-IV with granules throughout their length. Genital area, free sternites and anal operculum with few granules. Legs: (Figs 3A View Figure 3 , 7O, P View Figure 7 ) Coxae I-II each one with a retrolateral and prolateral apophyses. Coxa III with a prolateral apophysis. Coxa IV with setiferous granules distributed over the entire length. Trochanters I-III unarmed and with few granules. Trochanter IV with few granules and an apical retrolateral blunt tubercle. Femora I-III unarmed and with few granules. Femur IV with sparse granules; a retrolateral row of 15-17 acuminate tubercles over the entire length of the segment, except the base; a prolateral row of 10-12 acuminate tubercles along the distal ⅔; with four ventrodistal tubercles. Patellae I-III unarmed, with few granules. Patella IV with seven dorsal acuminate tubercles, the two largest apical ones; several granules on the ventral surface. Tibiae I-IV unarmed, with few granules. Tarsal formula: (n =3) 7-8 (8), 12-13 (1 3), 10-15 (10), 11-13 (13). Penis: (Fig. 12G, H View Figure 12 ) VP subrectangular, with distal margin straight and lateral projections; straight in lateral view. MS C1-C3 subapical long and straight; MS A1 median long and straight; MS B1 sub basal long and straight. Lateral sacs long, apically acuminate, with long T3-like microsetae. Stylus with a non-swollen apex and without apical projections. Dorsal process conical, with acuminated apex. Promontory straight. - FEMALE: Measurements (n =2) DSW: 4.2-4.5; DSL: 4.5-4.9; CL: 1.5-1.7. FIVL: 7.2-8.1. ChL: 1.7-1.8. Chelicerae not swollen. Femur IV with retrolateral row of 13-14 tubercles, smaller than those of the retrolateral row of the males; prolateral row and ventrodistal tubercles absent or in the form of granules. Patella IV with dorsodistal acuminate tubercles, but smaller than in males. Tarsal formula: (n =2) 7, 11-12, 9-10, 10-12.


It differs from other species of the genus by the combination of the following characteristics: Gamma-type DSS (Fig. 3A View Figure 3 ); area III of DS with a pair of large spines (also present in A. pasolinii sp. nov.); male femur IV armed (Fig. 7O, P View Figure 7 ); longer legs (ratio between FIVL and DSL greater than 1; 1.65 in A. spielbergi sp. nov.); and scutal areas light yellow, with series of dark spots (Fig. 22H View Figure 22 ).

Derivatio nominis.

The specific epithet of masculine gender, in the genitive form, dedicated to the American filmmaker, producer and screenwriter Steven Allen Spielberg.


(Fig. 28 View Figure 28 ) PERU. Cajamarca . Parque Nacional Cutervo.

Material examined.

Type material: Holotype ♂, ' PERU, Cajamarca , Parque Nacional Cutervo , Puente Suro, 6°12′10″S 78°44′22″W, 22/V/2010, R. Pinto-da-Rocha & D. Silva leg. (MUSM) GoogleMaps - Paratypes 2 ♂, 1 ♀, ‘ditto’ (MZSP 36995); Paratype 1 ♀, ' PERU, Cajamarca , Parque Nacional Cutervo , near to Cueva San Andreas, 22/V/2010, R. Pinto-da-Rocha & D. Silva leg. (MZSP 36996) . Additional material: 1 ♂, ' PERU, Cajamarca , Parque Nacional Cutervo , San Andreas de Cutervo, 13/VI/1996, S. Córdova leg. (MUSM 0501248) ; 1 ♀, ‘ditto’ (MUSM 0501247).