Nannaria stellaradix, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180 : 1

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scientific name

Nannaria stellaradix

sp. nov.

Nannaria stellaradix View in CoL sp. nov. Figs 42 View Figure 42 , 43 View Figure 43 , 44 Vernacular name: “Starroot’s Twisted-Claw Millipede" View Figure 44

Material examined.

Holotype: United States - Virginia • ♂; Montgomery County, Riner, 222 Milky way NW, in wetland below lake; 36.9662°N, - 80.4179°W; elev. 773 m; 17 Oct. 2014; hand collected; J. Means leg.; VTEC MPE002331. GoogleMaps

Paratype: United States - Virginia • 1 ♂; same collection data as holotype; VMNH MPE00241 View Materials GoogleMaps 3 ♀♀; same collection data as holotype; VTEC MPE00238 View Materials GoogleMaps -240 • 2 ♀♀; same collection data as holotype; VMNH MPE00242 View Materials GoogleMaps , 244.

Other material.

United States - Virginia • 1 ♂; Floyd County, 2 mi. SW of Copper Valley; 36.9702°N, - 80.5425°W; 15 Oct. 1974; R. Hoffman leg.; VMNH NAN0146 GoogleMaps 1 ♂; Floyd County, Big Indian Creek , ca. 2 mi. S of Copper Valley; 36.9615°N, - 80.5171°W; 16 Apr. 1970; B. Coombs leg.; VMNH NAN0147 GoogleMaps 1 ♂; Pulaski County, Powhatan Scout Reservation, ca. 1 mi. E. of Macks Creek P.O.; 36.9645°N, - 80.6626°W; 6 Mar. 1976; R. Hoffman leg.; VMNH NAN0165. For detailed collection data see Suppl. material 7 GoogleMaps .


Adult males of Nannaria stellaradix sp. nov. are distinct from other Nannaria and the sympatric N. wilsoni , based on the following combination of characters: Gonopods. Gonopodal acropodite continually curving medially before apex. Distal zone short, directed medially with small triangular lateral flange (Fig. 42A View Figure 42 , red arrow), not directed caudally with enveloping lateral and medial flanges as in N. hokie , or enlarged, laminate, as in N. wilsoni . Acropodite with small laminate medial flange (Fig. 42A View Figure 42 , red triangle). Basal zone> 1/3 length of acropodite, not <1/5 as in N. wilsoni . Prefemur with enlarged prefemoral spine. Prefemur with small, ventrally curving prefemoral process arising dorsomedially from spine, not from top of spine as in N. hokie , or long, medially curving, paralleling acropodite as in N. wilsoni . Color. Tergites with light orange paranotal spots (Fig. 43 View Figure 43 ). Dark brown background. Dorsum of collum smooth with orange and white margin.


♂ holotype (VTEC, MPE00233): BL = 23.8, CW = 2.9, IW = 1.8, ISW = 0.8, B11W = 3.7, B11H = 2.6; ♀ paratype (VTEC, MPE00238): BL = 25.4, CW = 3.4, IW = 2.2, ISW = 0.9, B11W = 3.9, B11H = 2.8.


Three populations from Pulaski and Floyd counties in Virginia (VMNH, NAN0146, 147, 165) show some morphological variation (Fig. 44 View Figure 44 ) that may indicate a separate species; however, without molecular evidence, specific recognition is not warranted. Notable variations in these populations include a prefemoral process arising from the top of the prefemoral spine (Fig. 44 View Figure 44 , red arrow) and the lack of an acropodite medial flange.


Known from a linear area in southern Virginia (Virginia: Floyd and Pulaski counties, Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: 11 km2; status: MRE.


Individuals of N. stellaradix sp. nov. were collected from mesic mixed hardwood and pine habitats predominately composed of maple and white pine; individuals were collected from under leaf litter.


This species is named after the owner of the property from which the holotype was found, an artist named Starroot. The specific name is a noun in apposition derived from the Latin stella -, star, and - radix, root.

Type locality.

United States, Virginia, Montgomery County, Riner, 222 Milky way NW, in wetland below lake, 36.9662°N, - 80.4179°W.