Ochthephilus kleebergi, Makranczy, 2014

Makranczy, György, 2014, Revision of the genus Ochthephilus Mulsant & Rey, 1856 (Coleoptera: Staphylinidae, Oxytelinae), Revue suisse de Zoologie 121 (4), pp. 457-694 : 620-622

Ochthephilus kleebergi sp. nov. Figs 486-488, 509-513

TYPE MATERIAL: HOLOTYPE (3): “ NEPAL: E-Nepal, Rolwaling Himal , Nyimare, 3300m, [27°54' N, 86°22' E], 19.V.2000, leg. A. Kleeberg [cloud forest; sifting leaf litter under wood, wet mosses]” ( ZMHB) . – PARATYPES (25): same data as holotype (coll. Kleeberg, 13, 3♀, 2, coll. Schülke, 13). – “ Rolwaling Himal , umg. Nyimare , 3300m, [27°54' N, 86°22' E], 19.V.2000, leg. A. Kleeberg [cloud forest; sifting leaf litter under wood, wet mosses]” (coll. Kleeberg, 1♀, HNHM, 13). – “Rolwaling Himal, Rolwaling Tal, zw. Simigaon u. Nyimare, 2600m, [*+27.90/+86.27*], 17.V.2000, leg. A. Kleeberg [cloud forest; sifting leaf litter at a small creek shore]” (coll. Kleeberg, 1♀, HNHM, 1♀), same but Rolwaling Ufer, 2700m [*+27.90/+86.28*] ( HNHM, 1♀). – “ Dolakha distr., SW Kalinchok Mt., 3100m [*+27.75/+86.03*], 19-23.IV.1995, leg. Martens & Schawaller (no. 421)” ( SMNS, 13) . – “ CHINA: YUNNAN: Dali Cangshan , ca. 2500m [*+25.60/+100.19*], 10.IV.2003, leg. G. de Rougemont, stream moss” (coll. Rougemont, OXUM, 23, 6, HNHM, 13, MHNG, 1, USNM, 1, MNHP, 1, MSNV, 1) .

DESCRIPTION: Measurements (n=10): HW = 0.59 (0.57-0.61); TW = 0.54 (0.52- 0.55); PW = 0.65 (0.62-0.67); SW = 0.91 (0.81-0.96); AW = 0.95 (0.80-1.04); HL = 0.46 (0.44-0.48); EL = 0.19 (0.18-0.21); TL = 0.10 (0.09-0.11); PL = 0.54 (0.52-0.57); SL = 1.09 (1.00-1.18); SC = 1.00 (0.90-1.08); FB = 2.18 (2.04-2.30); BL = 4.07 (3.50- 4.40) mm. Whole body very dark brown, head darkest, almost black, elytra sometimes very slightly lighter, distinctly reddish but still dark brown, scutellar area to shoulders, narrowly along suture and a broad transversal stripe at apex black. Antennae, mouthparts and legs dark brown, antenna blackish, tarsi and both ends of tibiae occasionally somewhat lighter. On lighter specimens legs reddish medium brown but mid-tibiae remain dark brown, blackish. Body with greasy lustre due to shallow and rather fine punctation on head and pronotum, but distinct coriaceous microsculpture on interspaces everywhere; elytral punctation fine, medium deep and dense, setation dense. Pubescence on elytra rather regularly spaced and strong, in contrast with the less conspicuous but much longer setation of head and pronotum: with fine and dense setae. Abdominal tergites with setae just as strong as elytral ones but much longer, especially at apices of tergites and adjacent to laterosternites. Head anteriad eyes and near inner posterior margin of eye with stronger and much longer bristles, as well as pronotal margin; at middle of tibiae with darker bristles. Elytral apex without conspicuous setae. Last tarsomere with a few setae only.

Forebody. Antenna as in Fig. 511. Clypeus (Fig. 509) almost impunctate (colliculate microsculptured), trapezoid, corners rounded, anterior edge gently arched; separated by impressed transversal line (frontoclypeal suture) across a shinier area. Supraantennal prominences well developed, feebly separated from clypeus/vertex by impressions. Vertex with oblique impressions in middle almost joining in V-shape. Temples (Fig. 510) slightly bulging, evenly curved, barely longer than half of eye length. Neck separated by an impressed transversal groove, microsculpture much stronger than on head, with transverse cells, no setation. Pronotum with a narrow marginal bead, getting inconspicuous anteriorly with pronotal corners strongly curved in ventral direction. Posterior pronotal angles well-formed, just slightly obtuse-angled, sides in posterior 1/2 insignificantly concave. 'Anchor' fully formed, longitudinal mid-

FIGS 431-436

Ochthephilus uhligi sp. n.; head and pronotum (431), side of head (432), antenna (433), elytra and base of abdomen (434), apices of tergites VII-VIII (435), apices of sternites VII-VIII (436). All SEM, dorsal views (except last, ventral). Scale bar = 0.23 mm for 432, 0.43 mm for 435-436, 0.5 mm for 431, 434, 0.62 mm for 433.

line as a slightly elevated, impunctate, weakly microsculptured line, parallel to this line two gentle, semi-longitudinal elongate elevations in anterior half of disc. In corners of anchor feeble, oblique impressions directed outwards, in middle at sides of midline two smaller impressions. Elytra (Fig. 512) slightly broadening posteriorly, sutural corners narrowly rounded; apical sides slightly oblique and in inner halves more or less straight. Elytral surface rather even with two shallow, very elongate impressions behind scutellum. Head with fine coriaceous/colliculate microsculpture, fading on elevated parts, stronger in impressions, on pronotum microsculpture slightly stronger and more even. Punctation on head fine, more dense on posterior part and sides, on pronotum more evenly spaced, average interspaces much larger than puncture diameters; elytral punctation more even and regularly spaced, average interspaces (with indistinct coriaceous microsculpture) about as puncture diameters, punctures discrete.

Abdomen. Compared to forebody, abdomen with much more sparse, finer, less distinct punctation, microsculpture on tergal apices fine coriaceous with moderately transverse cells. Tergite VII posterior margin with palisade fringe broadened in middle (Fig. 513) with more coarse spiniform processes. Tergite VIII basal edge evenly arched, without concavity in middle of basal sclerotized band; apical edge with sinuate (protruding) corners, and broad, moderately deep emargination in between. Sternite VIII with rounded apical corners, apex in males shallowly concave laterally, gently sinuate in middle; in females slightly more sinuate (convex) in middle. Tergite X unmodified, apex very slightly wider in males than in females. Aedeagus as in Fig. 486. Female ringstructures as in Figs 487-488.

ETYMOLOGY: Named after Andreas Kleeberg (Berlin, Germany), who participated in several expeditions to the Nepal Himalaya and greatly contributed to the knowledge to its fauna and was also the collector of the holotype specimen .

COMPARATIVE NOTES: Out of the large black species (outside the O. monticola complex) it shares the pronotal shape with O. ritae and O. tichomirovae (but different from O. szarukani ); the shorter antenna is similar to that of O. szarukani (but different from that of both O. ritae and O. tichomirovae ). From the remaining black species, O. zerchei , it is distinguishable by size.

DISTRIBUTION: Currently only known from Nepal and the Chinese province of

Yunnan.

BIONOMICS: Specimens were collected at rather high elevations, ca. 2600-3300 m (“cloud forest”) by sifting leaf litter under wood, wet mosses at banks of small mountain streams.