Hyperaspidini Mulsant, 1846

Biranvand, Amir, Tomaszewska, Wioletta, Nedvěd, Oldřich, Khormizi, Mehdi Zare, Nicolas, Vincent, Canepari, Claudio, Shakarami, Jahanshir, Fekrat, Lida & Fürsch, Helmut, 2017, Review of the tribe Hyperaspidini Mulsant (Coleoptera: Coccinellidae) from Iran, Zootaxa 4236 (2), pp. 311-326 : 313

Tribe Hyperaspidini Mulsant, 1846

Diagnosis. Hyperaspidini can be separated from other tribes of the subfamily Coccinellinae by the following combination of characters: body elongate oval, depressed to rounded, convex; dorsal surface glabrous (except in Blaisdelliana Gordon ); antenna short, composed of 9 to 11 antennomeres, club elongate, fusiform, terminal antennomere small, recessed in preceding antennomere, antennal insertion exposed or concealed; eye large, not emarginate by an eye canthus, finely faceted, without pubescence; maxillary palpus with apical palpomere securiform; scutellar shield usually large; elytral epipleuron narrow, usually excavated for reception of femoral apex (except Hyperaspidius Crotch and Blaisdelliana Gordon ); legs short, femur grooved or flattened for reception of tibia, protibia slender, unmodified, or expanded, tarsus cryptotetramerous; abdomen with 6 ventrites in both sexes; male genitalia with penis guide asymmetrical, toothed, penis guide rooted in phallobase; female genitalia with coxites usually short, transverse, styli reduced or absent ( Gordon 1985; Gordon & Canepari 2008).