Diplommatina Benson, 1849

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 203

Diplommatina Benson, 1849

Diplommatina BenSon, 1849: 193 . Type SpecieS (SubSequently deSignated by NeVill, 1878): Bulimus folliculus Pfeiffer, 1846 .

Diagnosis. Diplommatina is usually characterized by a prominent columellar lamella in the aperture, a parietal lamella inside the last whorl, a constriction near the beginning of the last whorl and usually smooth inner and outer surfaces of the operculum ( Fig. 5 ).

Remarks. There are exceptions with regard to all characters mentioned in the diagnosis of the genus. Among the species usually classified in Diplommatina , some species lack a columellar lamella in the aperture, a parietal lamella, or a constriction of the last whorl. We found an arcuate ridge on the inner surface of the operculum of Diplommatina javana ( Fig. 4 ), usually considered as a characteristic of Palaina .

Diplommatina in the traditional sense proved to be polyphyletic in molecular phylogenies ( Rundell 2008; Webster et al. 2012). Thus, the delimitation of this genus and its subdivisions has to be revised. This will also affect the delimitation of the former subgenera Sinica Möllendorff, 1885 and Pseudopalaina Möllendorff, 1897 , which have been used for species from Java. These are currently not in use, because the position and the strength of the folds in the last whorl, on which these taxa were based (see Kobelt 1902), are too variable to allow a clear subdivision of the genus (see also Vermeulen 1993). Note, that Pseudopalaina was first published by Möllendorff (1897) and included only Diplommatina (Pseudopalaina) sulcicollis Möllendorff, 1897 from Java. Thus, the designation of Palaina polymorpha Crosse, 1866 from Palau as type species of Pseudopalaina by Wenz (1939) is not valid and Pseudopalaina is not a synonym of Hungerfordia Beddome, 1889 (see Yamazaki et al. 2013), but would be available for a genus-level group including species from Java.

Currently, molecular data are not available from the type species of Diplommatina from the Himalaya nor of any species from Java classified as Diplommatina . The severely needed revision of the genus-level systematics of the Diplommatinidae is beyond the scope of this paper.

Figures

FIGURES 4 – 8. Opercula of Diplommatinidae. 4. Diplommatina javana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13261). 5. Diplommatina auriculata Möllendorff, 1897, inner Surface of operculum, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13258). 6. Palaina gedeana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13244). 7. Palaina nubigena Möllendorff, 1897, outer Surface of operculum, WeSt JaVa: Cianjur, Kandang Badak (ZMA 68468). 8. Arinia yanseni n. sp., inner Surface of operculum, EaSt JaVa: Rembang, Sumber PacuSen (MZB 18882). Scale bar = 0.07 mm for FigS 3 – 6, 0.1 mm for Fig. 7.