Trichophallus armatus, Ingrisch, 2024

Ingrisch, Sigfrid, 2024, Revision of the genus Trichophallus Ingrisch, 1998 with notes on the genera Secsiva Walker, 1869 and Subrioides C. Willemse, 1966 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 5442 (1), pp. 1-66 : 43-46

Trichophallus armatus sp. nov.

Fig. 21

Holotype (male): Indonesia: Papua, Paniai , 7.9.1939 (Naturalis Leiden).

Other specimens studied: New Guinea: New Guinea Exp. K.N.A.G. 1939, Araboebivak , 10 October 1939 ( Naturalis Leiden )— 1 male ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Dejateda at Mt. Barara , 11 September 1939 ( Naturalis Leiden )— 1 female ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 4 September 1939 ( Naturalis Leiden )— 1 female ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 7 September 1939 ( Naturalis Leiden )— 2 males (CW ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 17 September 1939 ( Naturalis Leiden )— 1 female ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 27 September 1939 ( Naturalis Leiden )— 1 male ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 29 September 1939 ( Naturalis Leiden )— 3 males, 1 female (CW ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 3° 50‘ S, 136° 15‘ E, 1 October 1939 ( Naturalis Leiden )— 1 female ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 17 October 1939 ( Naturalis Leiden )— 1 female ( Naturalis Leiden ); New Guinea Exp. K.N.A.G. 1939, Paniai , 19 October–20 November 1939 ( Naturalis Leiden )— 1 male ( Naturalis Leiden ); Irian Jaya , New Guinea (Neth.), Wisselmaren , Itouda , Kamo Valley , elev. 1500 m, 13 August 1955 (coll. J.L. Gressitt)— 1 male ( Bishop Museum , Honolulu ( BPBM)); New Guinea (Neth.), Wisselmaren, Moenemani, Kamo Valley , elev. 1500 m, 15 August 1962 (coll. J. Sedlacek)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Guinea (Neth.), Wisselmaren, Urapura, Kamo Valley , elev. 1530 m, 11 August 1955 (coll. J.L. Gressitt)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NW), Nabire, S. Geelvink Bay, elev. 0–30 m, 2–9 July 1962 (coll. J.L. Gressitt)— 3 males, 2 females (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NW), Wisselmaren, Enarotadi, elev. 1900–2000 m, 2–11 July 1962 (coll. J. Sedlacek)— 2 males, 2 females (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NW), Wisselmaren, Enarotadi, elev. 1800–1900 m, 20 July 1962 (coll. J. Sedlacek)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NW), Wisselmaren, Enarotadi, elev. 1850–1900 m, 28 July 1962 (coll. J. Sedlacek)— 1 female (Bishop Museum, Honolulu ( BPBM)); New Guinea ( NW), Wisselmaren, Enarotadi, elev. 1800–1900 m, 1–9 August 1962 (coll. J. Sedlacek)— 1 male (Bishop Museum, Honolulu ( BPBM)); New Guinea: New Guinea Exp. K.N.A.G. 1939, Araboebivak, 10 October 1939 (Naturalis Leiden)— 1 male (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Dejateda at Mt. Barara, 11 September 1939 (Naturalis Leiden)— 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 4 September 1939 (Naturalis Leiden)— 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 7 September 1939 (Naturalis Leiden)— 2 males (CW (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 17 September 1939 (Naturalis Leiden)— 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 27 September 1939 (Naturalis Leiden)— 1 male (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 29 September 1939 (Naturalis Leiden)— 3 males, 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 3°50‘S, 136°15‘E, 1 October 1939 (Naturalis Leiden)— 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 17 October 1939 (Naturalis Leiden)— 1 female (Naturalis Leiden); New Guinea Exp. K.N.A.G. 1939, Paniai, 19 October–20 November 1939 — 1 male (Naturalis Leiden) .

Diagnosis. Male cerci with a pair of narrow, parallel projections at end. Titillators with a pair of simple, at base out-curved sclerites and with a second pair of sclerites with angulate base and apex faintly sinuate before acute tip; laterally of both sclerites with a pair of membranous blades that carry long hairs or bristles along internal margin and have the apical-lateral margin recurved and darkened. Female subgenital plate with apical projections separated from basal area by a step.

Description. Small species. Tegmina surpassing hind knees (one female slightly brachypterous with tegmen not reaching hind knees). Femora with the following number of spines on ventral margins: fore femur 4–9 external, 3–7 internal; mid femur 5–7 external, 1–4 internal near base; hind femur 6–11 external, 0–8 internal (the latter very minute except for 1 or 2 near apex).

Hind knee lobes usually uni-spinose, but in several specimens single lobes (rarely all) are bi-spinose.

Male. Stridulatory file roughly 1.4 mm long with 128–133 teeth. Tenth abdominal tergite terminating into two obtuse lobes, wide and roundly excised in between. Epiproct triangularly rounded, dorsal surface grooved. Paraprocts with a small obtuse projection at ventral margin. Cerci elongate, little conical, and slightly curved; circa basal third with internal margin slightly widened and a tubercle at end of widened area; with two acute, apical-internal projections, the dorsal a little smaller than the ventral projection, both with a spinule at tip. Subgenital plate widely excised at apex, setose, especially at apical margin and dorsal (= internal) apical surface; styli small, little shorter than excised area.

Titillators separate, strongly curved in middle, apical part (behind curve) triangularly widened, apex subacute. With a pair of apical-lateral sclerites that are completely separated from titillators proper and connected with them by membranes, circa horse-shoe-shaped but with apical branch slightly hooked and acute, basal branch widened and subdivided into a truncate and in a rounded lobe; the membrane that is connecting the titillators with the apico-lateral sclerites widened toward base, forming a large lobe that is folded across thus forming two triangular lobes: the medial lobe carrying a curved row of long bristles, the lateral lobe includes a black triangular sclerite; apical angle of fold between both lobes with a short granular projection. Phallus membranes with a granular cap on both sides of apical-lateral sclerites.

Female. Tenth abdominal tergite divided in midline. Subgenital plate divided in midline by a narrow membranous suture; basal area narrow, separated by an oblique, curved step from apical lobes; connection between basal area and apical lobes sclerotised in medial, membranous in lateral areas; apical lobes rather short, broad, triangular with angle rounded and lateral-proximal angle angularly curved ventrad; basal-lateral areas greatly extended dorsad but not outlined against ventral area, narrow with obtuse-angular apical margin. Ovipositor rather short and stout.

Coloration. Yellowish brown with patterns 0-1-4. Pronotum with a medium brown medial band including pattern 3. Tegmen with light veins and dark cells. Variation: Vertex light but can be black in melanistic specimens. Patterns 2 and 4 missing in some individuals. Legs maculated in some individuals.

Measurements.—body: male 16–22, female 17–21; pronotum: male 5.0–5.3, female 4.8–5.3; tegmen: male 17.5–22.0, female 21–22; hind femur: male 12–14, female 13.5–15.0; longest antenna: female 60; ovipositor: 9–10 mm.

Etymology.—The name of the new species refers to the comparatively large and stout projections of the male titillators provided with acute tips; from Latin armatus = armed.

Figures

FIGURE 21. Trichophallus armatus sp. nov.: A–G, male: A, stridulatory file; B–D, titillators in ventro-apical (B), oblique ventro-apical (C), and oblique dorso-cranial view (D); E, abdominal apex in apical view; F, cercus; G, subgenital plate; H–K, female: H–J, subgenital plate in lateral (H, J) and in ventral (I) view; K, ovipositor.