Poa persica subsp. multiradiata (Trautv.) Soreng, Cabi & L.J. Gillespie, 2018

Gillespie, Lynn J., Soreng, Robert John, Cabi, Evren & Amiri, Neda, 2018, Phylogeny and taxonomic synopsis of Poa subgenus Pseudopoa (including Eremopoa and Lindbergella) (Poaceae, Poeae, Poinae), PhytoKeys 111, pp. 69-102 : 85-86

Poa persica subsp. multiradiata (Trautv.) Soreng, Cabi & L.J. Gillespie comb. nov. Fig. 3D, E

Poa palustris var. multiradiata Trautv., Trudy Imp. S.-Peterburgsk. Bot. Sada 4: 406. 1876. Poa multiradiata (Trautv.) Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 7: 620. 1880. Eremopoa multiradiata (Trautv.) Roshev., Fl. URSS 2: 430, t. 32. 1934. Eremopoa persica subsp. multiradiata (Trautv.) Tzvelev, Zlaki SSSR 479. 1976.

Nephelochloa tripolitana Boiss. & Blanche, Diagn. Pl. Orient., ser. 2, 4: 133-134. 1859. Poa persica var. major Boiss., Fl. Orient. 5: 610-611. 1884. Type protologue. Hab. ad margines semitarum inter hortos ad Tripolium Syriae (Blanche), circa Byrouth in Libano (Gaillardot). Type. Lebanon. S. Tripoli, dans les bords des chemins, 16 May 1854, Blanche 1267 (lectotype, here designated: JE [JE00005064 ex herb. Gaillardot, image!]). Note. Two of the original specimens turned up in our search, Blanche 1267 (JE00005064 ex herb. Gaillardot) and Gaillardot s.n. (JE00005065 ex herb Gaillardot no. 2323 [image!]). Blanche in 1869 (P02530724) might also be original material, with a distribution date rather than a collection date.

Eragrostis barbeyi Post, Bull. Herb. Boissier 5: 760-761. 1897. Type protologue. Habitat in collibus prope Midyat (Mardin), no. 38. Type. Turkey. Midyat, Hillsides, May 1895, 38 Barbey (lectotype, here designated by Nada Sinno Saoud & RJS: BEI! (image seen by RJS!)). Note. The BEI sheet has "No. 55 38 Barbey, 1895" (55 was originally written as 54 but the 4 written over by 5).

Eremopoa mardinensis R.R. Mill, Fl. Turkey & E. Aegean Isl. 9: 624, 488. 1985.Type. Turkey. Mardin, Mardin to Nusaybin, 8 km from Mardin, 850 m alt., shallow limestone gully, 22 May 1957, P. H. Davis & D. Hedge 28491 (holotype: E! [E00196494]).

Type.

Armenia rossica, prope monasterium Kiptschach, 1875, G. Raddi. Type: Armenia rossica: prope monasterium Kiptschach in monte Alagos, Jun 1875, G. Radde 124 (holotype: LE! [photo E00326521!]; isotypes: LE, LE, W [W19160014191 image!]).

Distribution.

Armenia, Georgia, Iran, Lebanon, Pakistan, Syria and Turkey.

Notes.

The presence of hairs on the lemmas in material treated as “multiradiat” is confused in the literature. Mill (1985) indicates that E. multiradiata and E. persica s.s. have lemma keels hairy in the lower ⅓- ½. We concur with Tzvelev (1976), who keyed E. persica subsp. persica as lemmas short pilose along the base of keel and marginal veins and subsp. multiradiata as lemmas glabrous or with a few solitary hairs.

Mill (1985) distinguished his new species Eremopoa mardinensis from E. multiradiata based on its glabrous lemmas, 8-12-flowered spikelets and florets strongly divergent from the rachilla. However, subsp. multiradiata also has glabrous lemmas (as noted above) and divergent florets (when spikelets are in flower) and its (4)5-9(10)-flowered spikelets overlap in number; therefore, we treat E. mardinensis as a synonym of E. multiradiata . The type material of Eragrostis barbeyi is from the same place as E. mardinensis and is clearly the same form (spikelets many-flowered); Nephelochloa tripolitana , with ca. 12-14-flowered spikelets, also appears to belong to this form. If E. mardinensis were accepted as a species, the basionym names Eragrostis barbeyi or Nephelochloa tripolitana would have priority.

Figures

Figure 3. Poa subgenus Pseudopoa sect. Pseudopoa. A P. diaphora subsp. diaphora var. diaphora, Chu, Kyrgyz Republic (Soreng et al. 7537) B, C P. persica subsp. persica, Adiyaman, Turkey (Soreng et al. 9215) B habit C closeup of base of plant showing keeled leaf sheaths and caniculate blades D, E P. persica subsp. multiradiata, Mardin, Turkey (Soreng et al. 9240) D habit E spikelet showing glabrous lemmas. Photos by R. J. Soreng.